Botanische Staatssammlung München © 1995–2005
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Taxon name:

Data maintenance

1. Data set source [mandatory] [mandatory character] 1. authentic 2. compiled 3. template 5. standard item
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2. Data set author(s) [mandatory] [mandatory character] () ()

3. Data set reviewer(s) [mandatory character] 23. Helms G. 40. Marbach B. 45. Nash T.H. 61. Sheard J.
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5. Data set publication permission [mandatory character] 1. no 2. yes None selected () ()

6. Data set publication date

Nomenclature: Taxonomic Hierarchy

7. Current taxonomic status 1. accepted 2. basionymous
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8. Taxonomic rank [mandatory] [mandatory character] 2. species 3. subspecies 4. variety 5. form None selected () ()

9. Currently accepted name

11. Genus name [mandatory] [mandatory character] () ()

12. Synonyms

13. Family name [mandatory] [mandatory character] 99. Physciaceae Zahlbr. (1898) None selected () ()

14. Suborder name 5. Lecanorineae None selected () ()

15. Order name 9. Lecanorales None selected () ()

Nomenclature: Type Information

16. Basionym

17. Type [mandatory character]

19. Protologue

20. Type specimen label data

Nomenclature: Taxonomic Literature

21. Taxonomic notes

22. Taxonomic literature

Biogeography

23. Hemisphere distribution () 1. Northern hemispheric ()

24. Floristic province {= zonal distribution} [mandatory] [mandatory character] 1. holarctic 2. arctic 3. subarctic 4. boreal 5. Mediterranean 6. subtropical 7. pantropical 8. neotropical 9. palaeotropic 10. subantarctic 11. antarctic 14. cosmopolitan 15. temperate
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25. Altitude [mandatory] [mandatory character] 1. coastal 2. colline 3. submontane 4. montane 5. upper montane 6. subalpine 7. alpine
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27. Continent distribution [mandatory] [mandatory character] 1. Africa 2. Antarctica 3. Asia-Temperate 4. Australasia 5. Europe 6. Northern America 7. Southern America 8. Pacific 9. Asia-Tropical
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38. Checklist records [mandatory] 1. Australia 2. Austria 7. Germany 10. Great Britain 11. Guianas 13. Italy 15. Mexico 17. New Guinea 18. New Zealand 20. Sonoran Desert 21. Sweden and Norway 22. Thailand 26. United States and Canada (continental)
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Ecology

43. Life habit () 2. biotroph (inhabiting living organisms) ()

44. Lifestyle [mandatory] [mandatory character] 4. lichenized (= parasitic and / or mutual symbiotic with photobionts) 5. lichenicolous (= parasitic on lichens)
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45. Substrate preference [mandatory character] 1. terricolous (on soil) 2. bryophytic (on bryophytes) 3. lignicolous (on wood) 4. corticolous (on bark) 6. saxicolous (on rock)
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46. Thallus-substrate interaction 1. endosubstratic 3. episubstratic
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47. Habitat insolation 1. sun-exposed 2. partially shaded 3. fully shaded
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48. Habitat moisture 1. dry 2. moist 3. wet
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53. Rock substrate mineral content 1. non-calciferous (lacking calcites) 2. calciferous (containing calcites)
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Host or Phorophyte Taxonomy

72. Host fungus or lichen genus 6. Aspicilia 17. Dimelaena 24. Lecanora 55. Rhizocarpon
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Thallus Morphology: General Morphology

104. Thallus growth form [mandatory character] 1. indistinct 2. crustose 3. foliose 4. fruticose
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105. Thallus primary subdivision 1. continuous 2. divided
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107. Thallus ramification pattern 1. isotomic-dichotomous 4. subdichotomous 5. irregular
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108. Crustose growth organization 1. leprose 2. granular 3. rimose 5. areolate (primarily areolate) 6. placodioid 7. squamulose 8. subfruticose (minutely fruticose, dactyliform)
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109. Crustose growth structure () 2. subsquamulose ()

110. Crustose growth aggregation () 2. scattered (dispersed) ()

111. Crustose growth continuity 1. continuous throughout 2. dispersed along the periphery, continuous in the centre 3. dispersed throughout and distinctly seperate
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112. Foliose growth organization () 1. lobed ()

113. Foliose lobe widening 1. widening 2. parallel
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114. Foliose lobe outline 1. elongate 2. lunate 3. linear 4. flabellate
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116. Foliose lobe orientation 1. horizontal 2. ascending 3. imbricate
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117. Foliose lobe front 1. angular 2. rounded
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118. Foliose lobe curvature 1. concave 2. subconcave 3. plane 4. subconvex 5. convex
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119. Fruticose growth organization 1. erect 4. subpendulous 5. pendulous
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121. Fruticose branch structure 1. terete 2. subterete 3. applanate (moderately flattened)
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122. Fruticose branch flattening 1. bifacial 2. unifacial
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126. Thallus parts imbrication 1. not imbricate 2. imbricate
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129. Thallus parts length [categorical] 1. short 2. long
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131. Thallus parts width [categorical] 1. narrow 2. wide
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132. Thallus parts width [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					mm wide

133. Thallus parts thickness [categorical] 1. thin 2. thick
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134. Thallus parts thickness [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					mm thick

Thallus Morphology: Thallus Size and Differentiation

151. Overall thallus diameter [categorical] 1. very small 2. small 3. medium-sized 4. large
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152. Overall thallus diameter [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					cm in diameter

156. Thallus apex width [categorical] 1. not distinctly broader than average 2. distinctly broader than average
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158. Thallus apex curvature 1. not curved 3. curved upwards 4. curved downwards
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159. Thallus apex outline 1. rounded 2. truncate
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Thallus Morphology: Outline

160. Thallus perimeter outline 1. irregular 2. orbicular
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161. Thallus perimeter zonation 1. simple 2. zonate 3. lobed 5. fimbriate
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162. Thallus perimeter delimitation 1. indistinct 2. distinct
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164. Thallus margin structure 3. dentate 4. laciniate 8. phyllidiate 11. lobulate 13. crenate
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165. Thallus margin orientation 1. down-turned 2. levelled 3. up-turned
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166. Thallus margin brightness [categorical] 1. paled 2. concolourous 3. darkened
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167. Primal thallus differentiation 1. peripheral (forming a prothallus) 2. extending (forming a hypothallus)
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168. Primal thallus persistence 1. disappearing 2. persistent
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170. Primal thallus colour 1. black 2. grey 6. brown 8. white
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Thallus Morphology: Upper Thallus Surface

172. Upper surface colour 2. white 3. pale grey (= greyish white, whitish grey) 4. grey 6. black 8. pale red (= reddish white, whitish red, incl. pink, rosé) 9. grey-red (= greyish red, reddish grey) 10. brownish red (= reddish brown) 15. grey-green (= greyish green, greenish gray) 16. brownish green (= greenish brown, incl. olive, olivaceous) 21. grey-blue (= bluish grey, greyish blue) 24. yellow 26. grey-yellow (= greyish yellow, yellowish grey) 27. brownish yellow (= yellowish brown) 30. brown 31. pale brown (= brownish white, whitish brown) 32. grey-brown (= greyish brown, brownish grey) 33. dark brown (= brownish black, blackish brown)
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175. Upper surface glossiness 1. matt (opaque, dull) 2. glossy (shiny) 3. vitreous
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176. Upper surface roughness 1. smooth 2. rough
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177. Upper surface texture 2. pulverulaceous 3. byssoid
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191. Upper surface maculae: presence 1. absent 2. present
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192. Upper surface maculae: outline 1. punctiform 2. orbicular 3. linear 4. irregular
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196. Upper surface pruina: presence 1. epruinose 2. pruinose
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197. Upper surface pruina: abundance 1. scarce 2. abundant
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198. Upper surface pruina: colour 2. greyish 4. greenish 8. whitish
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199. Upper surface pruina: distribution 1. continuous 2. patchy
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200. Upper surface pruina: location 1. central 3. peripheral
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201. Upper surface pseudocyphellae: presence 1. absent 2. present
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204. Upper surface pseudocyphellae: arrangement 1. isolated 2. reticulate
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208. Upper surface pseudocyphellae: location 1. central 2. laminal 3. peripheral
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210. Upper surface pseudocyphellae: position 1. submarginal 2. marginal
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218. Upper surface cilia: presence 1. eciliate 2. ciliate
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219. Upper surface cilia: kind 1. spinulous 2. slender
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225. Upper surface hairs: presence 1. absent 2. present
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227. Upper surface hairs: location 2. apical 3. laminal 4. interspersed 5. marginal
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228. Upper surface hairs position () 1. on thallus lobes or branches ()

230. Upper surface isidia: presence 1. absent 2. present
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231. Upper surface isidia: location () 2. laminal ()

232. Upper surface isidia: position {across the surface} 1. marginal 2. submarginal
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234. Upper surface isidia: arrangement 1. isolated 2. clustered
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235. Upper surface isidia: shape 1. globular 3. pustular 5. cylindrical 6. clavate 7. spathulate
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237. Upper surface isidia: ramification 1. not branched 2. branched
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238. Upper surface isidia: ramification pattern 1. sparse 2. coralloid 3. moderate
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240. Upper surface isidiomorphs: presence () 2. present ()

252. Upper surface soralia: presence 1. absent 2. present
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255. Upper surface soralia: colour 2. white 3. pale grey (= greyish white, whitish grey) 4. grey 10. brownish red (= reddish brown) 14. pale green (= greenish white, whitish green) 16. brownish green (= greenish brown, incl. olive, olivaceous) 20. pale blue (= whitish blue, bluish white) 24. yellow 25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine) 26. grey-yellow (= greyish yellow, yellowish grey) 27. brownish yellow (= yellowish brown) 30. brown 32. grey-brown (= greyish brown, brownish grey)
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256. Upper surface soralia: location 1. central 2. laminal 3. subperipheral 4. peripheral
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257. Upper surface soralia: position 1. apical 2. subapical 3. lateral
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258. Upper surface soralia: origin () 3. tubercular (eroding from the tip of tubercles, warts) ()

259. Upper surface soralia: shape 1. semicapitate 2. capitate 3. hooded (helmet-shaped) 4. spathulate 5. labriform 6. maculiform 7. ear-shaped
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260. Upper surface soralia: outline 1. erose (diffuse, unspecific) 2. punctiform 3. irregular ("effigurate") 4. linear
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261. Upper surface soralia: elevation () 3. prominent ()

262. Upper surface soralia: excavation 1. slightly excavated 2. fully excavated
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264. Upper surface soralia: differentiation 1. not confluent 2. confluent (but not fusing) 3. fusing
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268. Upper surface soralia: development {from isidia} () 2. isidiate ()

269. Upper surface soredia: diameter [categorical] 1. grainy (coarse) 2. farinose (fine)
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276. Upper surface blastidia: presence 1. absent 2. present
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277. Upper surface schizidia: presence 1. absent 2. present
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278. Upper surface lobules: presence 1. absent 2. present
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Thallus Morphology: Lower Thallus Surface

281. Lower surface presence 1. absent 2. present
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282. Lower surface overall colour 2. white 3. pale grey (= greyish white, whitish grey) 4. grey 6. black 10. brownish red (= reddish brown) 14. pale green (= greenish white, whitish green) 15. grey-green (= greyish green, greenish gray) 21. grey-blue (= bluish grey, greyish blue) 24. yellow 25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine) 26. grey-yellow (= greyish yellow, yellowish grey) 27. brownish yellow (= yellowish brown) 30. brown 31. pale brown (= brownish white, whitish brown)
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283. Lower surface central colour 2. white 4. grey 6. black 30. brown
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284. Lower surface peripheral colour 1. black 2. grey 6. brown 8. white 28. pale brown (= brownish white, whitish brown)
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285. Lower surface attachment extent 2. loosely attached 3. firmly attached
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286. Lower surface attachment kind 1. by the whole lower surface 2. by holdfasts 3. lacking
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296. Lower surface wrinkles: presence 1. absent 2. present
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297. Lower surface wrinkles: location 1. central 2. peripheral
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300. Lower surface rhizines: presence 1. absent 2. present (rhizinate)
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301. Lower surface rhizines: location 1. central 3. peripheral
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302. Lower surface rhizines: abundance 1. sparse 2. moderately abundant 3. abundant
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307. Lower surface rhizines: colour 2. white 3. pale grey (= greyish white, whitish grey) 4. grey 6. black 30. brown 31. pale brown (= brownish white, whitish brown)
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309. Lower surface rhizines: projections 1. absent 2. present
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310. Lower surface rhizines: ramification 1. not branched 2. branched
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311. Lower surface rhizines: ramification pattern 1. irregular 2. dichotomous 3. squarrose 4. sparsely furcate
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316. Lower surface soralia: presence 1. absent 2. present
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326. Lower surface tomentum: presence 1. absent 2. present
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Thallus Anatomy: Upper Cortex

329. Upper cortex: presence 1. absent 2. present
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331. Upper cortex height [percentage]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					% of thallus width

332. Upper cortex height [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm high

349. Upper cortex pigment location 1. in upper layer 2. in lower layer
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367. Upper cortex outer layer: structure 1. prosoplectenchymatous 2. paraplectenchymatous 3. scleroplectenchymatous
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371. Upper cortex inner layer: structure 1. prosoplectenchymatous 2. paraplectenchymatous 3. scleroplectenchymatous
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Thallus Anatomy: Photobiont Layer

375. Photobiont layer: zonation 1. single-layered 2. multi-layered
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Thallus Anatomy: Medulla

380. Medulla height [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm thick

383. Medulla colour 2. white 7. red 24. yellow 25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine) 34. orange (= reddish yellow, yellowish red)
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384. Medulla density 1. compact 2. dense 3. loose 4. very loose, becoming hollow
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388. Medulla hyphal arrangement 1. forming several bundles 2. forming a compact mass
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Thallus Anatomy: Lower Cortex

397. Lower cortex presence 1. absent 2. present
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400. Lower cortex height [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm thick

401. Lower cortex plectenchyma structure 1. paraplectenchymatous 2. prosoplectenchymatous
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404. Lower cortex hyphal orientation 1. anticlinal 2. periclinal
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Reproduction

414. Ascomata presence [mandatory] [mandatory character] 1. absent 2. present
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Ascocarp Morphology: General Ascocarp Morphology

418. Ascocarp structure [mandatory character] () 1. apothecioid ()

420. Ascocarp outline 1. orbicular 3. irregular
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421. Ascocarp location {on main thallus} 1. central 2. laminal 3. subperipheral 4. peripheral
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424. Ascocarp position {on primary thallus parts} 5. subterminal 6. terminal
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428. Ascocarp emergence () 1. not emerging (immersed, e.g., cryptolecanorine, cryptolecideine, aspicilioid) ()

429. Ascocarp attachment 1. adnate (semi-immersed) 2. sessile 3. substipitate
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430. Ascocarp basis 1. not constricted (flat below) 2. slightly constricted (moderately conical at the base) 3. strongly constricted (strongly conical at the base)
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431. Ascocarp abundance [categorical] 1. sparse 2. moderately abundant 3. abundant
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432. Ascocarp abundance [numerical]

Min:	Max:	Unit:
		per thallus

437. Ascocarp size [categorical] 1. small 2. large
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438. {orbicular} ascocarp diameter [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					mm in diam.

Ascocarp Morphology: Margin

447. Ascocarp margin: type 1. lecideine/biatorine 2. lecanorine (incl. zeorine)
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448. Ascocarp margin: structure 2. lobate 3. lobulate 4. crenulate 5. granulose 6. scabrose 7. verrucose 8. verruculose
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449. Ascocarp margin: roughness () 1. smooth ()

451. Ascocarp margin: persistency 1. persistent 2. excluded
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452. Ascocarp margin: development 1. indistinct (thin) 2. distinct (moderately thickened) 3. prominent (thick)
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457. Ascocarp margin: width [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm wide

460. Ascocarp margin: parathecial crown presence 1. absent 2. present
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461. Ascocarp margin: parathecial crown brightness [categorical] 1. darker than surrounding margin 2. concolourous with surrounding margin 3. brighter than surrounding margin
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463. Ascocarp margin: soredia 1. absent 2. present (sorediate)
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464. Ascocarp margin: isidia 1. absent 2. present (isidiate)
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466. Ascocarp margin: cilia 1. absent 2. present (ciliate)
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467. Ascocarp margin: hairs 1. absent 2. present (hirsute)
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Ascocarp Morphology: Disk

487. Ascocarp disk: curvature 1. urceolate (strongly concave) 2. excavate (concave) 3. plane (expanded) 4. weakly convex
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488. Ascocarp disk: colour 6. black 21. grey-blue (= bluish grey, greyish blue) 22. brownish blue (= bluish brown) 30. brown 32. grey-brown (= greyish brown, brownish grey)
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491. Ascocarp disk: pruina presence 1. absent 2. present
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492. Ascocarp disk: pruina abundance 1. scarce 2. abundant
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493. Ascocarp disk: pruina colour 1. white 2. yellow greenish 7. pale grey (= whitish grey, greyish white)
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Ascocarp Anatomy: Exciple

504. Ascocarp excipulum: pigment carbonization () 2. lecideine (carbonized) ()

505. Ascocarp excipulum: layer pigmentation 2. white 3. pale grey (= greyish white, whitish grey) 4. grey 5. dark gray (= greyish black, blackish grey) 6. black 7. red 8. pale red (= reddish white, whitish red, incl. pink, rosé) 9. grey-red (= greyish red, reddish grey) 10. brownish red (= reddish brown) 11. bluish red (= reddish blue, incl. violet, purple) 12. dark red (= blackish red, reddish black) 13. green 14. pale green (= greenish white, whitish green) 15. grey-green (= greyish green, greenish gray) 16. brownish green (= greenish brown, incl. olive, olivaceous) 17. bluish green (= greenish blue; incl. aeruginose, turquoise) 18. dark green (= blackish green, greenish black) 19. blue 20. pale blue (= whitish blue, bluish white) 21. grey-blue (= bluish grey, greyish blue) 22. brownish blue (= bluish brown) 23. dark blue (= blackish blue, bluish black) 24. yellow 25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine) 26. grey-yellow (= greyish yellow, yellowish grey) 27. brownish yellow (= yellowish brown) 28. greenish yellow (= yellowish green, incl. lime green) 29. dark yellow (= yellowish black, blackish yellow) 30. brown 31. pale brown (= brownish white, whitish brown) 32. grey-brown (= greyish brown, brownish grey) 33. dark brown (= brownish black, blackish brown) 34. orange (= reddish yellow, yellowish red)
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506. Ascocarp excipulum: crystal inspersion 1. not inspersed with crystals 2. inspersed with crystals
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Ascocarp Anatomy: Parathecium

507. Parathecium width [categorical] 1. narrow (indistinct, reduced) 2. broad (distinct)
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508. Parathecium width [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm wide

509. Parathecium structure 1. paraplectenchymatous 2. prosoplectenchymatous, strongly glutinized
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510. Parathecium pigmentation 2. white 3. pale grey (= greyish white, whitish grey) 4. grey 5. dark gray (= greyish black, blackish grey) 6. black 7. red 8. pale red (= reddish white, whitish red, incl. pink, rosé) 9. grey-red (= greyish red, reddish grey) 10. brownish red (= reddish brown) 11. bluish red (= reddish blue, incl. violet, purple) 12. dark red (= blackish red, reddish black) 13. green 14. pale green (= greenish white, whitish green) 15. grey-green (= greyish green, greenish gray) 16. brownish green (= greenish brown, incl. olive, olivaceous) 17. bluish green (= greenish blue; incl. aeruginose, turquoise) 18. dark green (= blackish green, greenish black) 19. blue 20. pale blue (= whitish blue, bluish white) 21. grey-blue (= bluish grey, greyish blue) 22. brownish blue (= bluish brown) 23. dark blue (= blackish blue, bluish black) 24. yellow 25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine) 26. grey-yellow (= greyish yellow, yellowish grey) 27. brownish yellow (= yellowish brown) 28. greenish yellow (= yellowish green, incl. lime green) 29. dark yellow (= yellowish black, blackish yellow) 30. brown 31. pale brown (= brownish white, whitish brown) 32. grey-brown (= greyish brown, brownish grey) 33. dark brown (= brownish black, blackish brown) 34. orange (= yellowish red, reddish yellow)
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514. Parathecium glutinization () 2. strongly glutinized ()

515. Parathecium crystal inspersion 1. not inspersed with crystals 2. inspersed with crystals
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516. Parathecium crystal solubility 1. dissolving in 10% KOH 2. not dissolving in 10% KOH
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Ascocarp Anatomy: General Amphithecium Anatomy

530. Amphithecium presence 1. absent 2. present
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538. Amphithecium height [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm high

540. Amphithecium width [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm wide

541. Amphithecium inspersion: type 1. not inspers 3. inspersed with crystals
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544. Amphithecium inspersion: crystal size () 1. small (< 5 µm in diameter) ()

545. Amphithecium inspersion: crystal abundance () 2. abundant ()

546. Amphithecium inspersion: crystal solubility 1. not dissolving in 10% KOH 2. dissolving in 10% KOH
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559. Amphithecial photobiont abundance 1. absent 2. sparse (eulecanorine) 3. abundant
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Ascocarp Anatomy: Lower Amphithecium

563. Lower amphithecium height [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm high

564. Lower amphithecium plectenchyma structure 1. pseudoparenchymatous 2. prosoplectenchymatous
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565. Lower amphithecium prosoplectenchyma hyphal orientation () 2. anticlinal ()

Ascocarp Anatomy: Amphithecial Cortex

566. Amphithecial cortex presence 1. absent 2. present
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572. Amphithecial cortex width [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm wide

574. Amphithecial cortex basal width [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm wide at the base

604. Amphithecial cortex: outer layer: pigmentation 2. white 3. pale grey (= greyish white, whitish grey) 4. grey 5. dark gray (= greyish black, blackish grey) 6. black 7. red 8. pale red (= reddish white, whitish red, incl. pink, rosé) 9. grey-red (= greyish red, reddish grey) 10. brownish red (= reddish brown) 11. bluish red (= reddish blue, incl. violet, purple) 12. dark red (= blackish red, reddish black) 13. green 14. pale green (= greenish white, whitish green) 15. grey-green (= greyish green, greenish gray) 16. brownish green (= greenish brown, incl. olive, olivaceous) 17. bluish green (= greenish blue; incl. aeruginose, turquoise) 18. dark green (= blackish green, greenish black) 19. blue 20. pale blue (= whitish blue, bluish white) 21. grey-blue (= bluish grey, greyish blue) 22. brownish blue (= bluish brown) 23. dark blue (= blackish blue, bluish black) 24. yellow 25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine) 26. grey-yellow (= greyish yellow, yellowish grey) 27. brownish yellow (= yellowish brown) 28. greenish yellow (= yellowish green, incl. lime green) 29. dark yellow (= yellowish black, blackish yellow) 30. brown 31. pale brown (= brownish white, whitish brown) 32. grey-brown (= greyish brown, brownish grey) 33. dark brown (= brownish black, blackish brown) 34. orange (= reddish yellow, yellowish red)
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608. Amphithecial cortex: inner layer: pigmentation 2. white 3. pale grey (= greyish white, whitish grey) 4. grey 5. dark gray (= greyish black, blackish grey) 6. black 7. red 8. pale red (= reddish white, whitish red, incl. pink, rosé) 9. grey-red (= greyish red, reddish grey) 10. brownish red (= reddish brown) 11. bluish red (= reddish blue, incl. violet, purple) 12. dark red (= blackish red, reddish black) 13. green 14. pale green (= greenish white, whitish green) 15. grey-green (= greyish green, greenish gray) 16. brownish green (= greenish brown, incl. olive, olivaceous) 17. bluish green (= greenish blue; incl. aeruginose, turquoise) 18. dark green (= blackish green, greenish black) 19. blue 20. pale blue (= whitish blue, bluish white) 21. grey-blue (= bluish grey, greyish blue) 22. brownish blue (= bluish brown) 23. dark blue (= blackish blue, bluish black) 24. yellow 25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine) 26. grey-yellow (= greyish yellow, yellowish grey) 27. brownish yellow (= yellowish brown) 28. greenish yellow (= yellowish green, incl. lime green) 29. dark yellow (= yellowish black, blackish yellow) 30. brown 31. pale brown (= brownish white, whitish brown) 32. grey-brown (= greyish brown, brownish grey) 33. dark brown (= brownish black, blackish brown) 34. orange (= reddish yellow, yellowish red)
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Ascocarp Anatomy: Episamma

622. Episamma presence 1. absent 2. present
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Ascocarp Anatomy: Epithecium

627. Epithecium presence 1. absent 2. present
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629. Epithecium height [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm high

630. Epithecial hyphae apical cell: shape 1. simple 2. spathulate 3. subulate
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639. Epithecial hyphae apical cell pigmentation {in H2O} 10. brownish red (= reddish brown) 16. brownish green (= greenish brown, incl. olive, olivaceous) 22. brownish blue (= bluish brown) 27. brownish yellow (= yellowish brown) 30. brown 31. pale brown (= brownish white, whitish brown)
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641. Epithecial hyphae apical cell: pigment solubility {in 10% KOH} 1. insoluble in 10% KOH 2. soluble in 10% KOH
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642. Epithecial hyphae apical cell: pigment reactivity () 2. 10% KOH + ()

Ascocarp Anatomy: Hymenium

653. Hymenium height [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm high

654. Hymenium pigmentation {in H2O} 2. white 25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine) 31. pale brown (= brownish white, whitish brown)
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659. Hymenium gel presence 1. absent 2. present (interascal hyphae gelatinized)
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661. Hymenium inspersion: type 1. not inspers 2. inspersed with oil droplets 3. inspersed with crystals
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667. Hymenium euamyloid reaction 1. Lugol’s negative 2. Lugol’s positive
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Ascocarp Anatomy: Interascal Hyphae

676. Interascal hyphae ramification 1. scarcely branched 2. distinctly branched
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677. Interascal hyphae anastomoses 1. not or scarcely anastomosed 2. distinctly anastomosed
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Ascocarp Anatomy: Subhymenium

684. Subhymenium height [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm high

699. Subhymenium pigmentation {in H2O} 2. white 3. pale grey (= greyish white, whitish grey) 4. grey 5. dark gray (= greyish black, blackish grey) 6. black 7. red 8. pale red (= reddish white, whitish red, incl. pink, rosé) 9. grey-red (= greyish red, reddish grey) 10. brownish red (= reddish brown) 11. bluish red (= reddish blue, incl. violet, purple) 12. dark red (= blackish red, reddish black) 13. green 14. pale green (= greenish white, whitish green) 15. grey-green (= greyish green, greenish gray) 16. brownish green (= greenish brown, incl. olive, olivaceous) 17. bluish green (= greenish blue; incl. aeruginose, turquoise) 18. dark green (= blackish green, greenish black) 19. blue 20. pale blue (= whitish blue, bluish white) 21. grey-blue (= bluish grey, greyish blue) 22. brownish blue (= bluish brown) 23. dark blue (= blackish blue, bluish black) 24. yellow 25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine) 26. grey-yellow (= greyish yellow, yellowish grey) 27. brownish yellow (= yellowish brown) 28. greenish yellow (= yellowish green, incl. lime green) 29. dark yellow (= yellowish black, blackish yellow) 30. brown 31. pale brown (= brownish white, whitish brown) 32. grey-brown (= greyish brown, brownish grey) 33. dark brown (= brownish black, blackish brown) 34. orange (= reddish yellow, yellowish red)
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Ascocarp Anatomy: Hypothecium

707. Hypothecium height [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm high

708. Hypothecium pigmentation {in H2O} 2. white 3. pale grey (= greyish white, whitish grey) 5. dark gray (= greyish black, blackish grey) 8. pale red (= reddish white, whitish red, incl. pink, rosé) 9. grey-red (= greyish red, reddish grey) 10. brownish red (= reddish brown) 14. pale green (= greenish white, whitish green) 15. grey-green (= greyish green, greenish gray) 16. brownish green (= greenish brown, incl. olive, olivaceous) 20. pale blue (= whitish blue, bluish white) 21. grey-blue (= bluish grey, greyish blue) 22. brownish blue (= bluish brown) 25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine) 26. grey-yellow (= greyish yellow, yellowish grey) 27. brownish yellow (= yellowish brown) 31. pale brown (= brownish white, whitish brown) 32. grey-brown (= greyish brown, brownish grey) 33. dark brown (= brownish black, blackish brown) 34. orange (= reddish yellow, yellowish red)
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716. Hypothecium prosoplectenchyma type 2. serioplectenchymatous 9. dense textura intricata
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717. Hypothecium hyphal orientation 2. anticlinal 3. irregular
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721. Hypothecium inspersion type 1. not inspers 2. inspersed with oil droplets
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Ascocarp Anatomy: Asci

751. Ascus shape 4. broadly clavate 5. clavate 6. narrowly clavate 7. obclavate 8. cylindrical
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753. Ascus length [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm long

754. Ascus width [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm wide

769. Ascus dehiscence type () 2. lecanoralean ()

Ascocarp Anatomy: Ascospores

782. Ascospore type 1. Beltraminia-type 2. Bicincta-type 3. Bischoffii-type 5. Callispora-type 7. Conradii-type 8. Diplotomma-type 9. Dirinaria-type 14. Milvina-type 15. Mischoblastia-type 16. Orcularia-type 17. Pachysporaria-type 19. Physcia-type 20. Physcia-type and Pachysporaria-type 21. Physconia-type 23. Rinodinella-type 27. Tunicata-type
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784. Ascospore number [categorical] 1. 1–2 per ascus 2. c. 4 per ascus 3. c. 8 per ascus 4. 12-16 per ascus 5. 16-32 per ascus 6. more than 32 per ascus
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785. Ascospore number [numerical]

Min:	Mean:	Upper range:	Max:	Unit:
				per ascus

786. Ascospore shape 3. oval 4. ellipsoid 5. broadly ellipsoid 16. fusiform
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787. Ascospore length [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm long

788. Ascospore width [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm wide

792. Ascospore septum presence 1. absent 2. present
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793. Ascospore septum orientation 1. transversely septate 2. muriform
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795. Ascospore {median} septum number [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					-transversally septate

799. Ascospore septum formation 1. early septum formation (Physciaceae-type A) 2. late septum formation (Physciaceae-type B)
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800. Ascospore cell lumen size [categorical] 1. even (all cells of equal size) 2. constricted in the centre (apically and basally enlarged) 3. tapered (swollen in the centre)
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802. Ascospore wall width [categorical] 1. thin 2. thick
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803. Ascospore wall width [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm wide

804. Ascospore wall differentiation {visible in the light microscope} () 2. distinctly differentiated (into primary and secondary wall) ()

805. {apical} ascospore wall thickening 1. absent 2. present
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806. {lateral} ascospore wall thickening 1. absent 2. present
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807. {septal} ascospore wall thickening 1. absent 2. presence
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809. Ascospore wall pigment development 1. remaining hyaline 2. becoming pigmented
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810. Ascospore wall pigmentation 2. grey 4. pale brown (= brownish white, whitish brown) 5. middle brown 7. dark brown (= brownish black, blackish brown)
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812. Ascospore perispore ornamentation 1. not ornamented (psilate, smooth) 2. ornamented
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813. Ascospore perispore surface structure 1. echinate 2. foveate 3. rugulate 4. striate 5. porate
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814. Ascospore mucilaginous sheath width [categorical] 1. indistinctly halonate (mucilaginous sheath absent or thin) 2. distinctly halonate (with a thick mucilaginous sheath)
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Conidioma Morphology

816. Conidiomata presence [mandatory character] 1. absent 2. present
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818. Conidiomata abundance 1. sparse 2. moderately abundant 3. strongly abundant
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Conidioma Anatomy: Conidiophores

857. Conidiophore type {sensu Vobis, Braun, etc.} 7. Vobis-type I 8. Vobis-type II 9. Vobis-type III 10. Vobis-type IV 11. Vobis-type V 12. Vobis-type VI 13. Vobis-type VII 14. Vobis-type VIII
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Conidioma Anatomy: Conidia

873. Conidium shape 1. globose 2. subglobose 3. oval 4. ellipsoid 5. broadly ellipsoid 11. bacilliform 13. cylindrical 16. fusiform
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879. Conidium length [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm long

880. Conidium width [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm wide

Chemistry: Secondary Metabolites

913. Known secondary metabolites: presence 1. absent 2. present
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914. Known secondary metabolites: location 2. only in the upper part of the cortex 4. throughout the thallus medulla 11. in soredia (or soralia)
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915. Known secondary metabolites: name 231. atranorin 250. chloroatranorin 336. gyrophoric acid 347. hopane-6a,22-diol 390. leucotylin 392. lichexanthone 442. miriquidic acid 470. norstictic acid 486. paludosic acid 491. pannarin 536. salazinic acid 537. scabrosin 4,4'-diacetate 538. scabrosin 4,4'-dibutyrate 539. scabrosin 4-acetate-4'-butyrate 540. scabrosin 4-acetate-4'-hexanoate 543. secalonic acid a 549. skyrin 559. stictic acid 588. variolaric acid
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916. Known secondary metabolites: substance class 2. (higher) aliphatic acids 3. (anthra-)quinones 13. secalonic acids
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917. Unknown secondary metabolites: presence 1. absent 2. present
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919. Unknown secondary metabolites: provisional name () 1. 'Heterodermia unknown spathulin' ()

921. Calcium oxalate crystals: presence 1. absent 2. present
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Chemistry: Spot Tests

926. Spot test reactions 1. absent 2. present
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927. Upper surface K reactivity 1. – (negative) 2. + deep yellow 5. + red
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928. Upper surface C reactivity 1. – 5. + red
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930. Upper surface KC reactivity 1. – 5. + red
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931. Upper surface PD reactivity 1. – 2. + yellow 8. + orange
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932. Cortex K reactivity 1. – 2. + deep yellow 3. + bright yellow (citrine, lemon)
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935. Cortex KC reactivity 1. – 2. + deep yellow 3. + bright yellow (citrine, lemon)
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936. Cortex PD reactivity 1. – 2. + yellow
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937. Medulla K reactivity 1. – 2. + yellow 3. + bright yellow (citrine, lemon) 4. + yellow changing to red 8. + orange
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938. Medulla C reactivity 1. – 5. + red
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940. Medulla KC reactivity 1. – 2. + yellow 3. + bright yellow (citrine, lemon) 5. + red 9. + orange
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941. Medulla PD reactivity 1. – 2. + deep yellow 5. + red
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942. Lower surface K reactivity 1. – 2. + deep yellow 5. + red
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947. Soredia K reactivity 1. – 2. + deep yellow 3. + pink 4. + orange
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949. Soredia KC reactivity 1. – 2. + deep yellow 3. + pink 4. + orange
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Chemistry: UV-Fluorescence

983. UV-fluorescence: upper surface 1. – (negative) 2. + (positive)
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