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Data Set Maintenance: Data set standard item. Data set author(s): Knoph J.-G., Rambold G. & Triebel D. (94-07-19). Data set reviewer(s): Kainz C. (00-11-15); to be published after submission; 94-07-19.

Nomenclature: Current taxonomic status: accepted. Taxonomic rank: species. Carbonea. Synonyms: Catillaria intrusa (Th.Fr.) Th.Fr., Lecidiopsis intrusa (Th.Fr.) Zopf, Conida intrusa (Th.Fr.) Sacc., Micarea intrusa (Th.Fr.) Coppins & Kilias, Lecidea aphanoides Nyl., Lecidea melaphana Nyl., Lecidea contrusa Vain., Lecidea halacsyi Steiner in Halacsy, Nesolechia halacsyi (Steiner) Steiner, Carbonea halacsyi (Steiner) Hafellner & Sancho, Nesolechia dispersula Rehm ex Sacc. & D. Sacc. in Sacc., Lecidea dispersula Arnold, Nesolechia dispersula Rehm ex Magnus in Dalla Torre & Sarnth.; Lecanoraceae Körb. (1855).

Taxonomic Literature: Knoph et al. in ed., Arnold (1899: 6 - sub Lecidea dispersula), Clauzade et al. (1989: 59 - sub Lecidea dispersula), Coppins (1983: 138-140 - sub Micarea intrusa), Hafellner & Sancho (1990: 365-366 - sub Carbonea halacsyi), Hertel (1970a: 424 - sub Lecidea intrudens, 1992: 300), Keissler (1930: 85-86 - sub Conida intrusa, 137 - sub Lecidea dispersula et Nesolechia dispersula), Magnusson (1946: 53 - sub Lecidea intrudens), Mus & Egea (1989: 120 - sub Micarea intrusa), Rambold & Triebel (1992), Wirth (1973: 186-187 - sub Catillaria intrusa), Zahlbruckner (1925: 513-514 - sub Lecidea aphanoides, 1925: 637, 1932a: 349 - sub Lecidea melaphana, 1927: 49 - sub Catillaria intrusa).

Ecology: Biotroph; lichenized or lichenicolous; growing on bark, non-calciferous rock, or calciferous rock; substrate non-calciferous. Host or Phorophyte Taxonomy: Amygdalaria, Aspicilia, Lecidea, Lecidella, Pertusaria, Rhizocarpon, Rinodina, and Tephromela, Hymeneliaceae, Lecanoraceae, Lecideaceae, Physciacaeae, and Pertusariaceae.

Thallus: Indistinct or crustose, rimose or areolate (primarily areolate); separate thallus parts thin (0.1 - 0.7 mm). Thallus Outline: Persistent. Upper Surface: Black, grey, or brown, rough; special structures absent. Lower Surface: Special structures present:

Reproduction Strategy: With sexual (and possible asexual) stages. Ascocarps: Soon sessile, slightly constricted at the base, .25-.45 mm in diam.. Margin: Excluded, indistinct to distinct. Disk: Plane to weakly convex, black, epruinose, without an umbo. Exciple: Green, brown, white, or brownish yellow. Parathecium (proper excipulum): Brown or white, dissolving in 10% KOH. Amphithecium (thalline excipulum): Present; 15 µm wide. Epithecium: Apical cells distinctly swollen, black or green. Hymenium: White (with orange patches). Hypothecium: White, brownish yellow, or orange.

Ascospores: Ellipsoid, oblong-obtuse, or oblong, (7)-7.1-10.5-14.4-(17) µm long, (3)-3.4-4.2-5.6-(6) µm wide; septa absent or present (occasionally).

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