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Data Set Maintenance: Data set compiled and standard item. Data set author(s): Kainz C. Data set reviewer(s): Schubert K. (06-03-23); revised.

Nomenclature: Current taxonomic status: accepted or basionymous. Taxonomic rank: species. Erysiphaceae Tul. & C. Tul.; Erysiphales.

Type Information: Basionym: Erysiphe howeana U. Braun. Type: Erysiphe howeana U. Braun.

Taxonomic Literature: Taxonomic notes: +appressoria moderately lobed;+conidiophores foot-cells straight, cylindric, followed by 1-2 shorter cells or cells of about the same length;+ascocarp outer wall cells obscure, irregularly polygonal, 8-25 µm diam.;. Braun U., Beih. Nova Hedwigia 89: 1-700 [204-205] (1987); Braun U., The powdery mildews (Erysiphales) of Europe. - 1-337. Jena, Stuttgart, New York (1995).

Biogeography: Continent: Africa, Europe, Northern America (USA, throughout the entire range of the hosts; Canada; introduced in Europe and South Africa), and Southern America (Argentina). Country or state(s): United Kingdom, Sweden, Germany, Hungary, Poland, Switzerland (incl. Liechtenstein), France (excl. Corsica), Portugal, Spain (incl. Andorra & Monaco), Italy (incl. San Marino & The Vatican City, excl. Sicily, Sardinia), Romania, European Turkey, Belarus, Baltic States (Estonia, Lithuania, Latvia), and Ukraine; Quebec (Canada).

Ecology: Biotroph; phytopathogenic; growing on leaves, amphigenous (sometimes covering the entire surface of the leaves). Host or Phorophyte Taxonomy: Oenothera sp.; Oenothera, Onagraceae.

Reproduction Strategy: With sexual (and possible asexual) stages. Ascocarps: Cleistothecioid, orbicular, forming independently from the host thallus or mycelium, scattered or in loose groups, (.075)-.09-.14-(.16) mm in diam.. Margin: External filaments present; interlaced with each other or interwoven with surrounding mycelium, mycelioid, straight, .5-2 µm long, 4-9 µm in diameter, hyaline or pigmented (in the lower half, yellowish to brownish, paler upwards), numerous, growing all across the lower half of the ascocarp, thin, not ramified, septate.

Asci: 3-15 asci per ascocarp, not stipitate or indistinctly stipitate, 40-80 µm long, 24-40 µm wide; dehiscence unitunicate.

Ascospores: c. 4 to c. 8 per ascus, spores 3-5-6 per ascus, ellipsoid or ovoid, (16)-18-23-(24) µm long, 9-14.5 µm wide; septa absent.

Conidiophores: Pseudoidium-type; not branched; basal cells (20)-25-35-(45) µm long, 7.5-11 µm wide. Conidium Formation: Conidiogenous cells single. Conidia: Ellipsoid, ovoid, doliiform, or sub-cylindrical; macroconidial (germ tubes arising from an end, moderately long, terminating with an unlobed or lobed appressorium, ca. 3.5-5 µm wide), not branched, 24-35-(43) µm long, 14-21-(22.5) µm wide; aseptate.

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