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Go to: Data maintenance    Nomenclature: Taxonomic Hierarchy    Nomenclature: Type Information    Nomenclature: Taxonomic Literature    Biogeography:    Ecology:    Host or Phorophyte Taxonomy:    Lichen Photobionts:    Thallus Morphology: General Morphology    Thallus Morphology: Thallus Differentiation    Thallus Morphology: Outline    Thallus Morphology: Upper Thallus Surface    Thallus Morphology: Lower Thallus Surface    Thallus Anatomy: Upper Cortex    Thallus Anatomy: Medulla    Thallus Anatomy: Lower Cortex    Ascocarp Morphology: General Ascocarp Morphology    Ascocarp Morphology: Wall    Ascocarp Morphology: Margin    Ascocarp Anatomy: Exciple    Ascocarp Anatomy: Amphithecial Cortex    Ascocarp Anatomy: Periphyses    Ascocarp Anatomy: Episamma    Ascocarp Anatomy: Epithecium    Ascocarp Anatomy: Hymenium    Ascocarp Anatomy: Interascal Hyphae    Ascocarp Anatomy: Asci    Ascocarp Anatomy: Ascospores    Conidioma Morphology    Conidioma Anatomy: Conidia    Chemistry: Secondary Metabolites    Chemistry: Spot Tests    Index of all items of the project


Taxon name:



Data maintenance   back to top

1. Data set source [mandatory] [mandatory character]     2. compiled     3. template     5. standard item
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2. Data set author(s) [mandatory] [mandatory character]     8. Breuss O.     28. Kainz C.     42. Nash T.H.     60. Triebel D.  None selected () ()
3. Data set reviewer(s) [mandatory character]     28. Kainz C.     45. Nash T.H.
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4. Data set revision status [mandatory character]     1. not revised     2. revised  None selected () ()

Nomenclature: Taxonomic Hierarchy   back to top

7. Current taxonomic status    ()  1. accepted ()
8. Taxonomic rank [mandatory] [mandatory character]     2. species     3. subspecies     4. variety     5. form  None selected () ()
10. Taxon number
Mean:

11. Genus name [mandatory] [mandatory character]     218. Endococcus     438. Muellerella     810. Verrucaria  None selected () ()
12. Synonyms


13. Family [mandatory] [mandatory character]     139. Verrucariaceae Zenker (1827)  None selected () ()
15. Order [mandatory] [mandatory character]     22. Verrucariales  None selected () ()

Nomenclature: Type Information   back to top

16. Basionym


17. Type [mandatory character]


19. Protologue


20. Type specimen label data



Nomenclature: Taxonomic Literature   back to top

21. Taxonomic literature


22. Taxonomic notes



Biogeography:   back to top

23. Hemisphere distribution    ()  1. Northern hemispheric ()
24. Floristic province {= zonal distribution} [mandatory] [mandatory character]     1. holarctic     2. arctic     3. subarctic     4. boreal     5. Mediterranean     6. subtropical     7. pantropical     8. neotropical     9. palaeotropic     10. subantarctic     11. antarctic     14. cosmopolitan
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25. Altitude [mandatory] [mandatory character]     1. coastal     2. colline     3. submontane     4. montane     5. upper montane     6. subalpine     7. alpine
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27. Continent distribution [mandatory] [mandatory character]     1. Africa     2. Antarctica     3. Asia-Temperate     4. Australasia     5. Europe     6. Northern America     7. Southern America     8. Pacific     9. Asia-Tropical
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38. Checklist records [mandatory] [mandatory character]     15. Mexico     20. Sonoran Desert     26. United States and Canada (continental)
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Ecology:   back to top

43. Life habit     1. saprobic (= destruents)     2. biotroph (inhabiting living organisms)
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44. Lifestyle [mandatory] [mandatory character]     4. lichenized (= parasitic and / or mutual symbiotic with photobionts)     5. lichenicolous (= parasitic on lichens)
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45. Substrate preference [mandatory character]     1. terricolous (on soil)     2. bryophytic (on bryophytes)     3. lignicolous (on wood)     4. corticolous (on bark)     6. saxicolous (on rock)
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46. Thallus-substrate interaction     1. endosubstratic     3. episubstratic
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47. Habitat insolation     1. sun-exposed     2. partially shaded     3. fully shaded
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48. Habitat moisture     1. dry     2. moist     3. wet
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53. Rock substrate mineral content     1. non-calciferous (lacking calcites)     2. calciferous (containing calcites)
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Host or Phorophyte Taxonomy:   back to top

71. Host fungus species


72. Host fungus genus     1. Acarospora     2. Amandinea     3. Amygdalaria     4. Arthonia     5. Arthothelium     6. Aspicilia     7. Baeomyces     8. Bellemerea     9. Biatora     10. Buellia     11. Candelariella     12. Caloplaca     13. Cladonia     14. Cryptothecia     15. Dactylina     16. Dibaeis     17. Dimelaena     18. Diploicia     19. Diploschistes     20. Echinoplaca     21. Endocarpon     22. Graphis     23. Haematomma     24. Lecanora     25. Lecidea     26. Lecidella     27. Lepraria     28. Leprocaulon     29. Lobaria     30. Mazosia     31. Nephroma     32. Ochrolechia     33. Orphniospora     34. Pannaria     35. Parmelia     36. Parmentaria     37. Peltigera     38. Pertusaria     39. Phaeophyscia     40. Phaeorrhiza     41. Phlyctis     42. Phyllophiale     43. Physcia     44. Pilophorus     45. Placopsis     46. Pleopsidium     47. Polyblastia     48. Porina     49. Porpidia     50. Protoparmelia     51. Pseudocyphellaria     52. Psoroma     53. Pyrenula     54. Ramalina     55. Rhizocarpon     56. Rhizoplaca     57. Rinodina     58. Roccella     59. Solorina     60. Sporastatia     61. Sporopodium     62. Stereocaulon     63. Sticta     64. Tephromela     65. Thelotrema     66. Trapelia     67. Trichothelium     68. Umbilicaria     69. Xanthoria
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Lichen Photobionts:   back to top

80. Primary photobiont kind    ()  2. chlorococcal ()
86. Primary photobiont genus     15. Coccobotrys     20. Dilabifilum     26. Heterococcus     32. Myrmecia     43. Stichococcus
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Thallus Morphology: General Morphology   back to top

104. Thallus growth form [mandatory character]     1. indistinct     2. crustose     3. foliose
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105. Thallus primary subdivision     1. continuous     2. divided
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108. Crustose growth organization     2. granular     5. areolate (primarily areolate)     6. placodioid     7. squamulose
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109. Crustose growth structure    ()  2. subsquamulose ()
110. Crustose growth aggregation     1. adjoining (not dispersed)     2. scattered (dispersed)
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112. Foliose growth organization    ()  2. umbilicate ()
117. Foliose lobe front     1. angular     2. rounded
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132. Thallus parts width [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
mm wide

133. Thallus parts thickness [categorical]     1. thin     2. thick
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134. Thallus parts thickness [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
mm thick


Thallus Morphology: Thallus Differentiation   back to top

151. Overall thallus diameter [categorical]     1. very small     2. small     3. medium-sized     4. large
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152. Overall thallus diameter [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
cm in diameter

154. Thallus joints main branch {basal} constriction     1. basally not constricted     2. basally constricted
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Thallus Morphology: Outline   back to top

160. Thallus perimeter outline     1. irregular     2. orbicular
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161. Thallus perimeter zonation     1. simple     2. zonate     3. lobed     4. arachnoid
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162. Thallus perimeter delimitation     1. indistinct     2. distinct
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164. Thallus margin structure     1. entire     6. undulate     9. crenulate     11. lobulate
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Thallus Morphology: Upper Thallus Surface   back to top

171. Upper surface aspect    ()  1. verrucose ()
172. Upper surface colour     4. grey     10. brownish red (= reddish brown)     16. brownish green (= greenish brown, incl. olive, olivaceous)     27. brownish yellow (= yellowish brown)     30. brown     32. grey-brown (= greyish brown, brownish grey)     33. dark brown (= brownish black, blackish brown)
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175. Upper surface glossiness     1. matt (opaque, dull)     2. glossy (shiny)     3. vitreous
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176. Upper surface roughness     1. smooth     2. rough
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177. Upper surface texture     3. byssoid     4. erose
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179. Upper surface fissures: presence     1. absent     2. present
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181. Upper surface fissures: development     1. sparingly developed     2. strongly developed
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196. Upper surface pruina: presence     1. epruinose     2. pruinose
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197. Upper surface pruina: abundance     1. scarce     2. abundant
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198. Upper surface pruina: colour     2. greyish     8. whitish
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199. Upper surface pruina: distribution     1. continuous     2. patchy
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Thallus Morphology: Lower Thallus Surface   back to top

281. Lower surface presence     1. absent     2. present
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282. Lower surface overall colour     6. black     10. brownish red (= reddish brown)     27. brownish yellow (= yellowish brown)     31. pale brown (= brownish white, whitish brown)     32. grey-brown (= greyish brown, brownish grey)     33. dark brown (= brownish black, blackish brown)
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286. Lower surface attachment kind     1. by the whole lower surface     2. by holdfasts
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296. Lower surface wrinkles: presence     1. absent     2. present
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298. Lower surface wrinkles: arrangement     1. isolated     2. reticulate
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300. Lower surface rhizines: presence     1. absent     2. present (rhizinate)
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302. Lower surface rhizines: abundance     1. sparse     2. moderately abundant     3. abundant
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307. Lower surface rhizines: colour     10. brownish red (= reddish brown)     27. brownish yellow (= yellowish brown)
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Thallus Anatomy: Upper Cortex   back to top

329. Upper cortex: presence     1. absent     2. present
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332. Upper cortex height [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm high

334. Upper cortex plectenchyma structure     1. prosoplectenchymatous     2. paraplectenchymatous
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338. Upper cortex layer layer differentiation     1. one     2. two
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367. Upper cortex outer layer: structure     1. prosoplectenchymatous     2. paraplectenchymatous
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371. Upper cortex inner layer: structure     1. prosoplectenchymatous     2. paraplectenchymatous
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Thallus Anatomy: Medulla   back to top

380. Medulla height [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm thick

383. Medulla colour     2. white     31. pale brown (= brownish white, whitish brown)
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384. Medulla density     1. compact     2. dense     3. loose     4. very loose, becoming hollow
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Thallus Anatomy: Lower Cortex   back to top

397. Lower cortex presence     1. absent     2. present
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400. Lower cortex height [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm thick


Ascocarp Morphology: General Ascocarp Morphology   back to top

418. Ascocarp structure [mandatory character]    ()  2. perithecioid ()
428. Ascocarp emergence    ()  1. not emerging (immersed, e.g., cryptolecanorine, cryptolecideine, aspicilioid) ()
429. Ascocarp attachment     1. adnate (semi-immersed)     2. sessile
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438. {orbicular} ascocarp diameter [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
mm in diam.


Ascocarp Morphology: Wall   back to top

443. Ascocarp wall: carbonization     1. not carbonized     2. carbonized only in parts     3. carbonized around the ostiole, lacking a clypeus     4. with a carbonized black clypeus around the ostiole
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444. Ascocarp wall: fusion {of neighboured perithecia}     1. absent     2. present
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Ascocarp Morphology: Margin   back to top

453. Ascocarp margin: colour     6. black     25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine)     27. brownish yellow (= yellowish brown)     31. pale brown (= brownish white, whitish brown)
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Ascocarp Anatomy: Exciple   back to top

506. Ascocarp exciple: crystal inspersion    ()  1. not inspersed with crystals ()

Ascocarp Anatomy: Amphithecial Cortex   back to top

604. Amphithecial cortex: outer layer: pigmentation     2. white     6. black     27. brownish yellow (= yellowish brown)
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Ascocarp Anatomy: Periphyses   back to top

613. Periphyses presence     1. absent     2. present
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621. Periphyses gelatinization     1. absent     2. present (with mucilage between the periphyses)
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Ascocarp Anatomy: Episamma   back to top

622. Episamma presence     1. absent     2. present
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623. Episamma structure    ()  1. crystalline ()

Ascocarp Anatomy: Epithecium   back to top

627. Epithecium presence     1. absent     2. present
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639. Epithecial hyphae apical cell pigmentation {in H2O}     1. hyaline (translucent)     25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine)     27. brownish yellow (= yellowish brown)
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Ascocarp Anatomy: Hymenium   back to top

659. Hymenium gel presence     1. absent     2. present (interascal hyphae gelatinized)
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667. Hymenium euamyloid reaction     1. Lugol’s negative     2. Lugol’s positive
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Ascocarp Anatomy: Interascal Hyphae   back to top

673. Interascal hyphae presence     1. absent     2. present
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675. Interascal hyphae orientation     1. irregular     2. anticlinal
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679. Interascal hyphae cell length [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm long


Ascocarp Anatomy: Asci   back to top

751. Ascus shape     3. sacculiform     4. broadly clavate     5. clavate     6. narrowly clavate     8. cylindrical
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753. Ascus length [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm long

754. Ascus width [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm wide

769. Ascus dehiscence type    ()  4. verrucarioid ()

Ascocarp Anatomy: Ascospores   back to top

783. Ascospore arrangement     1. not uniseriate     2. uniseriate
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784. Ascospore number [categorical]     1. 1-2 per ascus     2. c. 4 per ascus     3. c. 8 per ascus     4. 12-16 per ascus
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785. Ascospore number [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
per ascus

786. Ascospore shape     2. subglobose     3. oval     4. ellipsoid     5. broadly ellipsoid     16. fusiform
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787. Ascospore length [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm long

788. Ascospore width [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm wide

792. Ascospore septum presence     1. absent     2. present
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793. Ascospore septum orientation     1. transversely septate     2. muriform
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795. Ascospore {median} septum number [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
-transversally septate

796. Ascospore {longitudinal} septum number
Min: Lower range: Mean: Upper range: Max: Unit:
longitudinal septa per transversal septum

802. Ascospore wall width [categorical]     1. thin     2. thick
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803. Ascospore wall width [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm wide

804. Ascospore wall differentiation {visible in the light microscope}    ()  2. distinctly differentiated (into primary and secondary wall) ()
810. Ascospore wall pigmentation     1. hyaline (translucent)     2. grey     4. pale brown (= brownish white, whitish brown)     5. middle brown     7. dark brown (= brownish black, blackish brown)
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812. Ascospore perispore ornamentation     1. not ornamented (psilate, smooth)     2. ornamented
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Conidioma Morphology   back to top

816. Conidiomata presence [mandatory character]     1. absent     2. present
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(Select a character state: to add this note/comment: )
817. Conidiomata kind    ()  1. pycnidial ()
818. Conidiomata abundance     1. sparse     2. moderately abundant     3. strongly abundant
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )

Conidioma Anatomy: Conidia   back to top

873. Conidium shape     10. oblong-obtuse     11. bacilliform
(Select a character state: to add this modifier: )
(Select a character state: to add this note/comment: )
879. Conidium length [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm long

880. Conidium width [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm wide


Chemistry: Secondary Metabolites   back to top

913. Known secondary metabolites: presence    ()  1. absent ()

Chemistry: Spot Tests   back to top

926. Spot test reactions    ()  1. absent ()

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Index of all items of the project

In case that additional characters and states are required to be included in this data set, consult the LIAS Instructions to Participants and follow the procedures described there.

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