Botanische Staatssammlung München © 1995–2004
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Taxon name:

Data maintenance

1. Data set source [mandatory] [mandatory character] 2. compiled 3. template 5. standard item
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2. Data set author(s) [mandatory] [mandatory character] 8. Breuss O. 28. Kainz C. 42. Nash T.H. 60. Triebel D. None selected () ()

3. Data set reviewer(s) [mandatory character] 28. Kainz C. 45. Nash T.H.
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4. Data set revision status [mandatory character] 1. not revised 2. revised None selected () ()

Nomenclature: Taxonomic Hierarchy

7. Current taxonomic status () 1. accepted ()

8. Taxonomic rank [mandatory] [mandatory character] 2. species 3. subspecies 4. variety 5. form None selected () ()

10. Taxon number

Mean:

11. Genus name [mandatory] [mandatory character] 218. Endococcus 438. Muellerella 810. Verrucaria None selected () ()

12. Synonyms

13. Family [mandatory] [mandatory character] 139. Verrucariaceae Zenker (1827) None selected () ()

15. Order [mandatory] [mandatory character] 22. Verrucariales None selected () ()

Nomenclature: Type Information

16. Basionym

17. Type [mandatory character]

19. Protologue

20. Type specimen label data

Nomenclature: Taxonomic Literature

21. Taxonomic literature

22. Taxonomic notes

Biogeography:

23. Hemisphere distribution () 1. Northern hemispheric ()

24. Floristic province {= zonal distribution} [mandatory] [mandatory character] 1. holarctic 2. arctic 3. subarctic 4. boreal 5. Mediterranean 6. subtropical 7. pantropical 8. neotropical 9. palaeotropic 10. subantarctic 11. antarctic 14. cosmopolitan
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25. Altitude [mandatory] [mandatory character] 1. coastal 2. colline 3. submontane 4. montane 5. upper montane 6. subalpine 7. alpine
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27. Continent distribution [mandatory] [mandatory character] 1. Africa 2. Antarctica 3. Asia-Temperate 4. Australasia 5. Europe 6. Northern America 7. Southern America 8. Pacific 9. Asia-Tropical
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38. Checklist records [mandatory] [mandatory character] 15. Mexico 20. Sonoran Desert 26. United States and Canada (continental)
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Ecology:

43. Life habit 1. saprobic (= destruents) 2. biotroph (inhabiting living organisms)
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44. Lifestyle [mandatory] [mandatory character] 4. lichenized (= parasitic and / or mutual symbiotic with photobionts) 5. lichenicolous (= parasitic on lichens)
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45. Substrate preference [mandatory character] 1. terricolous (on soil) 2. bryophytic (on bryophytes) 3. lignicolous (on wood) 4. corticolous (on bark) 6. saxicolous (on rock)
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46. Thallus-substrate interaction 1. endosubstratic 3. episubstratic
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47. Habitat insolation 1. sun-exposed 2. partially shaded 3. fully shaded
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48. Habitat moisture 1. dry 2. moist 3. wet
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53. Rock substrate mineral content 1. non-calciferous (lacking calcites) 2. calciferous (containing calcites)
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Host or Phorophyte Taxonomy:

71. Host fungus species

72. Host fungus genus 1. Acarospora 2. Amandinea 3. Amygdalaria 4. Arthonia 5. Arthothelium 6. Aspicilia 7. Baeomyces 8. Bellemerea 9. Biatora 10. Buellia 11. Candelariella 12. Caloplaca 13. Cladonia 14. Cryptothecia 15. Dactylina 16. Dibaeis 17. Dimelaena 18. Diploicia 19. Diploschistes 20. Echinoplaca 21. Endocarpon 22. Graphis 23. Haematomma 24. Lecanora 25. Lecidea 26. Lecidella 27. Lepraria 28. Leprocaulon 29. Lobaria 30. Mazosia 31. Nephroma 32. Ochrolechia 33. Orphniospora 34. Pannaria 35. Parmelia 36. Parmentaria 37. Peltigera 38. Pertusaria 39. Phaeophyscia 40. Phaeorrhiza 41. Phlyctis 42. Phyllophiale 43. Physcia 44. Pilophorus 45. Placopsis 46. Pleopsidium 47. Polyblastia 48. Porina 49. Porpidia 50. Protoparmelia 51. Pseudocyphellaria 52. Psoroma 53. Pyrenula 54. Ramalina 55. Rhizocarpon 56. Rhizoplaca 57. Rinodina 58. Roccella 59. Solorina 60. Sporastatia 61. Sporopodium 62. Stereocaulon 63. Sticta 64. Tephromela 65. Thelotrema 66. Trapelia 67. Trichothelium 68. Umbilicaria 69. Xanthoria
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Lichen Photobionts:

80. Primary photobiont kind () 2. chlorococcal ()

86. Primary photobiont genus 15. Coccobotrys 20. Dilabifilum 26. Heterococcus 32. Myrmecia 43. Stichococcus
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Thallus Morphology: General Morphology

104. Thallus growth form [mandatory character] 1. indistinct 2. crustose 3. foliose
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105. Thallus primary subdivision 1. continuous 2. divided
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108. Crustose growth organization 2. granular 5. areolate (primarily areolate) 6. placodioid 7. squamulose
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109. Crustose growth structure () 2. subsquamulose ()

110. Crustose growth aggregation 1. adjoining (not dispersed) 2. scattered (dispersed)
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112. Foliose growth organization () 2. umbilicate ()

117. Foliose lobe front 1. angular 2. rounded
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132. Thallus parts width [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					mm wide

133. Thallus parts thickness [categorical] 1. thin 2. thick
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134. Thallus parts thickness [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					mm thick

Thallus Morphology: Thallus Differentiation

151. Overall thallus diameter [categorical] 1. very small 2. small 3. medium-sized 4. large
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152. Overall thallus diameter [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					cm in diameter

154. Thallus joints main branch {basal} constriction 1. basally not constricted 2. basally constricted
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Thallus Morphology: Outline

160. Thallus perimeter outline 1. irregular 2. orbicular
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161. Thallus perimeter zonation 1. simple 2. zonate 3. lobed 4. arachnoid
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162. Thallus perimeter delimitation 1. indistinct 2. distinct
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164. Thallus margin structure 1. entire 6. undulate 9. crenulate 11. lobulate
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Thallus Morphology: Upper Thallus Surface

171. Upper surface aspect () 1. verrucose ()

172. Upper surface colour 4. grey 10. brownish red (= reddish brown) 16. brownish green (= greenish brown, incl. olive, olivaceous) 27. brownish yellow (= yellowish brown) 30. brown 32. grey-brown (= greyish brown, brownish grey) 33. dark brown (= brownish black, blackish brown)
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175. Upper surface glossiness 1. matt (opaque, dull) 2. glossy (shiny) 3. vitreous
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176. Upper surface roughness 1. smooth 2. rough
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177. Upper surface texture 3. byssoid 4. erose
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179. Upper surface fissures: presence 1. absent 2. present
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181. Upper surface fissures: development 1. sparingly developed 2. strongly developed
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196. Upper surface pruina: presence 1. epruinose 2. pruinose
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197. Upper surface pruina: abundance 1. scarce 2. abundant
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198. Upper surface pruina: colour 2. greyish 8. whitish
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199. Upper surface pruina: distribution 1. continuous 2. patchy
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Thallus Morphology: Lower Thallus Surface

281. Lower surface presence 1. absent 2. present
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282. Lower surface overall colour 6. black 10. brownish red (= reddish brown) 27. brownish yellow (= yellowish brown) 31. pale brown (= brownish white, whitish brown) 32. grey-brown (= greyish brown, brownish grey) 33. dark brown (= brownish black, blackish brown)
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286. Lower surface attachment kind 1. by the whole lower surface 2. by holdfasts
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296. Lower surface wrinkles: presence 1. absent 2. present
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298. Lower surface wrinkles: arrangement 1. isolated 2. reticulate
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300. Lower surface rhizines: presence 1. absent 2. present (rhizinate)
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302. Lower surface rhizines: abundance 1. sparse 2. moderately abundant 3. abundant
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307. Lower surface rhizines: colour 10. brownish red (= reddish brown) 27. brownish yellow (= yellowish brown)
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Thallus Anatomy: Upper Cortex

329. Upper cortex: presence 1. absent 2. present
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332. Upper cortex height [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm high

334. Upper cortex plectenchyma structure 1. prosoplectenchymatous 2. paraplectenchymatous
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338. Upper cortex layer layer differentiation 1. one 2. two
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367. Upper cortex outer layer: structure 1. prosoplectenchymatous 2. paraplectenchymatous
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371. Upper cortex inner layer: structure 1. prosoplectenchymatous 2. paraplectenchymatous
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Thallus Anatomy: Medulla

380. Medulla height [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm thick

383. Medulla colour 2. white 31. pale brown (= brownish white, whitish brown)
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384. Medulla density 1. compact 2. dense 3. loose 4. very loose, becoming hollow
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Thallus Anatomy: Lower Cortex

397. Lower cortex presence 1. absent 2. present
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400. Lower cortex height [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm thick

Ascocarp Morphology: General Ascocarp Morphology

418. Ascocarp structure [mandatory character] () 2. perithecioid ()

428. Ascocarp emergence () 1. not emerging (immersed, e.g., cryptolecanorine, cryptolecideine, aspicilioid) ()

429. Ascocarp attachment 1. adnate (semi-immersed) 2. sessile
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438. {orbicular} ascocarp diameter [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					mm in diam.

Ascocarp Morphology: Wall

443. Ascocarp wall: carbonization 1. not carbonized 2. carbonized only in parts 3. carbonized around the ostiole, lacking a clypeus 4. with a carbonized black clypeus around the ostiole
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444. Ascocarp wall: fusion {of neighboured perithecia} 1. absent 2. present
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Ascocarp Morphology: Margin

453. Ascocarp margin: colour 6. black 25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine) 27. brownish yellow (= yellowish brown) 31. pale brown (= brownish white, whitish brown)
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Ascocarp Anatomy: Exciple

506. Ascocarp exciple: crystal inspersion () 1. not inspersed with crystals ()

Ascocarp Anatomy: Amphithecial Cortex

604. Amphithecial cortex: outer layer: pigmentation 2. white 6. black 27. brownish yellow (= yellowish brown)
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Ascocarp Anatomy: Periphyses

613. Periphyses presence 1. absent 2. present
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621. Periphyses gelatinization 1. absent 2. present (with mucilage between the periphyses)
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Ascocarp Anatomy: Episamma

622. Episamma presence 1. absent 2. present
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623. Episamma structure () 1. crystalline ()

Ascocarp Anatomy: Epithecium

627. Epithecium presence 1. absent 2. present
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639. Epithecial hyphae apical cell pigmentation {in H2O} 1. hyaline (translucent) 25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine) 27. brownish yellow (= yellowish brown)
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Ascocarp Anatomy: Hymenium

659. Hymenium gel presence 1. absent 2. present (interascal hyphae gelatinized)
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667. Hymenium euamyloid reaction 1. Lugol’s negative 2. Lugol’s positive
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Ascocarp Anatomy: Interascal Hyphae

673. Interascal hyphae presence 1. absent 2. present
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675. Interascal hyphae orientation 1. irregular 2. anticlinal
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679. Interascal hyphae cell length [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm long

Ascocarp Anatomy: Asci

751. Ascus shape 3. sacculiform 4. broadly clavate 5. clavate 6. narrowly clavate 8. cylindrical
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753. Ascus length [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm long

754. Ascus width [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm wide

769. Ascus dehiscence type () 4. verrucarioid ()

Ascocarp Anatomy: Ascospores

783. Ascospore arrangement 1. not uniseriate 2. uniseriate
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784. Ascospore number [categorical] 1. 1-2 per ascus 2. c. 4 per ascus 3. c. 8 per ascus 4. 12-16 per ascus
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785. Ascospore number [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					per ascus

786. Ascospore shape 2. subglobose 3. oval 4. ellipsoid 5. broadly ellipsoid 16. fusiform
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787. Ascospore length [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm long

788. Ascospore width [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm wide

792. Ascospore septum presence 1. absent 2. present
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793. Ascospore septum orientation 1. transversely septate 2. muriform
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795. Ascospore {median} septum number [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					-transversally septate

796. Ascospore {longitudinal} septum number

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					longitudinal septa per transversal septum

802. Ascospore wall width [categorical] 1. thin 2. thick
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803. Ascospore wall width [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm wide

804. Ascospore wall differentiation {visible in the light microscope} () 2. distinctly differentiated (into primary and secondary wall) ()

810. Ascospore wall pigmentation 1. hyaline (translucent) 2. grey 4. pale brown (= brownish white, whitish brown) 5. middle brown 7. dark brown (= brownish black, blackish brown)
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812. Ascospore perispore ornamentation 1. not ornamented (psilate, smooth) 2. ornamented
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Conidioma Morphology

816. Conidiomata presence [mandatory character] 1. absent 2. present
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817. Conidiomata kind () 1. pycnidial ()

818. Conidiomata abundance 1. sparse 2. moderately abundant 3. strongly abundant
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Conidioma Anatomy: Conidia

873. Conidium shape 10. oblong-obtuse 11. bacilliform
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879. Conidium length [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm long

880. Conidium width [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm wide

Chemistry: Secondary Metabolites

913. Known secondary metabolites: presence () 1. absent ()

Chemistry: Spot Tests

926. Spot test reactions () 1. absent ()

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