Taxon name:

Data maintenance  

1. Data set source [mandatory] [mandatory character]     2. compiled ()     3. template     5. standard item
(Select a character state: 2 3 5 to add this modifier: )
(Select a character state: 2 3 5 to add this note/comment: )

2. Data set author(s) [mandatory] [mandatory character]     8. Breuss O.     28. Kainz C.     42. Nash T.H.     60. Triebel D.  None selected () ()

3. Data set reviewer(s)     28. Kainz C.     45. Nash T.H.
(Select a character state: 28 45 to add this modifier: )
(Select a character state: 28 45 to add this note/comment: )

4. Data set revision status [mandatory character]     1. not revised     2. revised  None selected () ()

Nomenclature: Taxonomic Hierarchy  

7. Current taxonomic status    ()  1. accepted ()

8. Taxonomic rank [mandatory] [mandatory character]     2. species     3. subspecies     4. variety     5. form  None selected () ()

10. Taxon number

11. Genus name [mandatory] [mandatory character]     218. Endococcus     438. Muellerella     810. Verrucaria  None selected () ()

12. Synonyms

13. Family name [mandatory] [mandatory character]     139. Verrucariaceae Zenker (1827)  None selected () ()

15. Order name     22. Verrucariales  None selected () ()

Nomenclature: Type Information  

16. Basionym


17. Type


19. Protologue


20. Type specimen label data



Nomenclature: Taxonomic Literature  

21. Taxonomic notes


22. Taxonomic literature

Biogeography  

23. Hemisphere distribution    ()  1. Northern hemispheric ()

24. Floristic province {= zonal distribution} [mandatory] [mandatory character]     1. holarctic ()     2. arctic     3. subarctic     4. boreal     5. Mediterranean     6. subtropical     7. pantropical     8. neotropical     9. palaeotropic     10. subantarctic     11. antarctic     14. cosmopolitan
(Select a character state: 1 2 3 4 5 6 7 8 9 10 11 14 to add this modifier: )
(Select a character state: 1 2 3 4 5 6 7 8 9 10 11 14 to add this note/comment: )

25. Altitude [mandatory] [mandatory character]     1. coastal     2. colline     3. submontane     4. montane     5. upper montane     6. subalpine     7. alpine
(Select a character state: 1 2 3 4 5 6 7 to add this modifier: )
(Select a character state: 1 2 3 4 5 6 7 to add this note/comment: )

27. Continent distribution [mandatory] [mandatory character]     1. Africa     2. Antarctica     3. Asia-Temperate     4. Australasia     5. Europe     6. Northern America     7. Southern America     8. Pacific     9. Asia-Tropical
(Select a character state: 1 2 3 4 5 6 7 8 9 to add this modifier: )
(Select a character state: 1 2 3 4 5 6 7 8 9 to add this note/comment: )

38. Checklist records [mandatory] [mandatory character]     15. Mexico     20. Sonoran Desert     26. United States and Canada (continental) ()
(Select a character state: 15 20 26 to add this modifier: )
(Select a character state: 15 20 26 to add this note/comment: )

Ecology  

43. Life habit     1. saprobic (= destruents)     2. biotroph (inhabiting living organisms)
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

44. Lifestyle [mandatory] [mandatory character]     4. lichenized (= parasitic and / or mutual symbiotic with photobionts)     5. lichenicolous (= parasitic on lichens)
(Select a character state: 4 5 to add this modifier: )
(Select a character state: 4 5 to add this note/comment: )

45. Substrate preference [mandatory character]     1. terricolous (on soil)     2. bryophytic (on bryophytes)     3. lignicolous (on wood)     4. corticolous (on bark)     6. saxicolous (on rock)
(Select a character state: 1 2 3 4 6 to add this modifier: )
(Select a character state: 1 2 3 4 6 to add this note/comment: )

46. Thallus-substrate interaction     1. endosubstratic     3. episubstratic
(Select a character state: 1 3 to add this modifier: )
(Select a character state: 1 3 to add this note/comment: )

47. Habitat insolation     1. sun-exposed     2. partially shaded     3. fully shaded
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

48. Habitat moisture     1. dry     2. moist     3. wet
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

53. Rock substrate mineral content     1. non-calciferous (lacking calcites)     2. calciferous (containing calcites)
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

Host or Phorophyte Taxonomy  

71. Host fungus or lichen species

72. Host fungus or lichen genus     1. Acarospora     2. Amandinea     3. Amygdalaria     4. Arthonia     5. Arthothelium     6. Aspicilia     7. Baeomyces     8. Bellemerea     9. Biatora     10. Buellia     11. Candelariella     12. Caloplaca     13. Cladonia     14. Cryptothecia     15. Dactylina     16. Dibaeis     17. Dimelaena     18. Diploicia     19. Diploschistes     20. Echinoplaca     21. Endocarpon     22. Graphis     23. Haematomma     24. Lecanora     25. Lecidea     26. Lecidella     27. Lepraria     28. Leprocaulon     29. Lobaria     30. Mazosia     31. Nephroma     32. Ochrolechia     33. Orphniospora     34. Pannaria     35. Parmelia     36. Parmentaria     37. Peltigera     38. Pertusaria     39. Phaeophyscia     40. Phaeorrhiza     41. Phlyctis     42. Phyllophiale     43. Physcia     44. Pilophorus     45. Placopsis     46. Pleopsidium     47. Polyblastia     48. Porina     49. Porpidia     50. Protoparmelia     51. Pseudocyphellaria     52. Psoroma     53. Pyrenula     54. Ramalina     55. Rhizocarpon     56. Rhizoplaca     57. Rinodina     58. Roccella     59. Solorina     60. Sporastatia     61. Sporopodium     62. Stereocaulon     63. Sticta     64. Tephromela     65. Thelotrema     66. Trapelia     67. Trichothelium     68. Umbilicaria     69. Xanthoria
(Select a character state: 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 43 44 45 46 47 48 49 50 51 52 53 54 55 56 57 58 59 60 61 62 63 64 65 66 67 68 69 to add this modifier: )
(Select a character state: 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 43 44 45 46 47 48 49 50 51 52 53 54 55 56 57 58 59 60 61 62 63 64 65 66 67 68 69 to add this note/comment: )

Lichen Photobionts  

80. Primary photobiont kind    ()  2. chlorococcal ()

86. Primary photobiont genus     15. Coccobotrys     20. Dilabifilum     26. Heterococcus     32. Myrmecia     43. Stichococcus
(Select a character state: 15 20 26 32 43 to add this modifier: )
(Select a character state: 15 20 26 32 43 to add this note/comment: )

Thallus Morphology: General Morphology  

104. Thallus growth form [mandatory character]     1. indistinct     2. crustose     3. foliose
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

105. Thallus primary subdivision     1. continuous     2. divided
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

108. Crustose growth organization     2. granular     5. areolate (primarily areolate)     6. placodioid     7. squamulose
(Select a character state: 2 5 6 7 to add this modifier: )
(Select a character state: 2 5 6 7 to add this note/comment: )

109. Crustose growth structure     2. subsquamulose     4. peltate
(Select a character state: 2 4 to add this modifier: )
(Select a character state: 2 4 to add this note/comment: )

110. Crustose growth aggregation     1. adjoining (not dispersed)     2. scattered (dispersed)
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

112. Foliose growth organization    ()  2. umbilicate ()

117. Foliose lobe front     1. angular     2. rounded
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

132. Thallus parts width [numerical]

133. Thallus parts thickness [categorical]     1. thin     2. thick
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

134. Thallus parts thickness [numerical]

Thallus Morphology: Thallus Size and Differentiation  

151. Overall thallus diameter [categorical]     1. very small     2. small     3. medium-sized     4. large
(Select a character state: 1 2 3 4 to add this modifier: )
(Select a character state: 1 2 3 4 to add this note/comment: )

152. Overall thallus diameter [numerical]

154. Thallus joints main branch {basal} constriction     1. basally not constricted     2. basally constricted
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

Thallus Morphology: Outline  

160. Thallus perimeter outline     1. irregular     2. orbicular
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

161. Thallus perimeter zonation     1. simple     2. zonate     3. lobed     4. arachnoid
(Select a character state: 1 2 3 4 to add this modifier: )
(Select a character state: 1 2 3 4 to add this note/comment: )

162. Thallus perimeter delimitation     1. indistinct     2. distinct
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

164. Thallus margin structure     1. entire     6. undulate     9. crenulate     11. lobulate
(Select a character state: 1 6 9 11 to add this modifier: )
(Select a character state: 1 6 9 11 to add this note/comment: )

Thallus Morphology: Upper Thallus Surface  

171. Upper surface aspect    ()  1. verrucose ()

172. Upper surface colour     4. grey     10. brownish red (= reddish brown)     16. brownish green (= greenish brown, incl. olive, olivaceous)     27. brownish yellow (= yellowish brown)     30. brown     32. grey-brown (= greyish brown, brownish grey)     33. dark brown (= brownish black, blackish brown)
(Select a character state: 4 10 16 27 30 32 33 to add this modifier: )
(Select a character state: 4 10 16 27 30 32 33 to add this note/comment: )

175. Upper surface glossiness     1. matt (opaque, dull)     2. glossy (shiny)     3. vitreous
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

176. Upper surface roughness     1. smooth     2. rough
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

177. Upper surface texture     3. byssoid     4. erose
(Select a character state: 3 4 to add this modifier: )
(Select a character state: 3 4 to add this note/comment: )

179. Upper surface fissures: presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

181. Upper surface fissures: development     1. sparingly developed     2. strongly developed
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

196. Upper surface pruina: presence     1. epruinose     2. pruinose
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

197. Upper surface pruina: abundance     1. scarce     2. abundant
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

198. Upper surface pruina: colour     2. greyish     8. whitish
(Select a character state: 2 8 to add this modifier: )
(Select a character state: 2 8 to add this note/comment: )

199. Upper surface pruina: distribution     1. continuous     2. patchy
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

Thallus Morphology: Lower Thallus Surface  

281. Lower surface presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

282. Lower surface overall colour     6. black     10. brownish red (= reddish brown)     27. brownish yellow (= yellowish brown)     31. pale brown (= brownish white, whitish brown)     32. grey-brown (= greyish brown, brownish grey)     33. dark brown (= brownish black, blackish brown)
(Select a character state: 6 10 27 31 32 33 to add this modifier: )
(Select a character state: 6 10 27 31 32 33 to add this note/comment: )

286. Lower surface attachment kind     1. by the whole lower surface     2. by holdfasts
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

296. Lower surface wrinkles: presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

298. Lower surface wrinkles: arrangement     1. isolated     2. reticulate
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

300. Lower surface rhizines: presence     1. absent     2. present (rhizinate)
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

302. Lower surface rhizines: abundance     1. sparse     2. moderately abundant     3. abundant
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

307. Lower surface rhizines: colour     10. brownish red (= reddish brown)     27. brownish yellow (= yellowish brown)
(Select a character state: 10 27 to add this modifier: )
(Select a character state: 10 27 to add this note/comment: )

Thallus Anatomy: Upper Cortex  

329. Upper cortex: presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

332. Upper cortex height [numerical]

334. Upper cortex plectenchyma structure     1. prosoplectenchymatous     2. paraplectenchymatous
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

338. Upper cortex layer layer differentiation     1. one     2. two
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

367. Upper cortex outer layer: structure     1. prosoplectenchymatous     2. paraplectenchymatous
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

371. Upper cortex inner layer: structure     1. prosoplectenchymatous     2. paraplectenchymatous
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

Thallus Anatomy: Medulla  

380. Medulla height [numerical]

383. Medulla colour     2. white     31. pale brown (= brownish white, whitish brown)
(Select a character state: 2 31 to add this modifier: )
(Select a character state: 2 31 to add this note/comment: )

384. Medulla density     1. compact     2. dense     3. loose     4. very loose, becoming hollow
(Select a character state: 1 2 3 4 to add this modifier: )
(Select a character state: 1 2 3 4 to add this note/comment: )

Thallus Anatomy: Lower Cortex  

397. Lower cortex presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

400. Lower cortex height [numerical]

Ascocarp Morphology: General Ascocarp Morphology  

418. Ascocarp structure [mandatory character]    ()  2. perithecioid ()

428. Ascocarp emergence    ()  1. not emerging (immersed, e.g., cryptolecanorine, cryptolecideine, aspicilioid) ()

429. Ascocarp attachment     1. adnate (semi-immersed)     2. sessile
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

438. {orbicular} ascocarp diameter [numerical]

Ascocarp Morphology: Wall  

443. Ascocarp wall: carbonization     1. not carbonized     2. carbonized only in parts     3. carbonized around the ostiole, lacking a clypeus     4. with a carbonized black clypeus around the ostiole
(Select a character state: 1 2 3 4 to add this modifier: )
(Select a character state: 1 2 3 4 to add this note/comment: )

444. Ascocarp wall: fusion {of neighboured perithecia}     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

Ascocarp Morphology: Margin  

453. Ascocarp margin: colour     6. black     25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine)     27. brownish yellow (= yellowish brown)     31. pale brown (= brownish white, whitish brown)
(Select a character state: 6 25 27 31 to add this modifier: )
(Select a character state: 6 25 27 31 to add this note/comment: )

Ascocarp Anatomy: Exciple  

506. Ascocarp excipulum: crystal inspersion    ()  1. not inspersed with crystals ()

Ascocarp Anatomy: Amphithecial Cortex  

604. Amphithecial cortex: outer layer: pigmentation     2. white     6. black     27. brownish yellow (= yellowish brown)
(Select a character state: 2 6 27 to add this modifier: )
(Select a character state: 2 6 27 to add this note/comment: )

Ascocarp Anatomy: Periphyses  

613. Periphyses presence     1. absent     2. present ()
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

621. Periphyses gelatinization     1. absent     2. present (with mucilage between the periphyses)
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

Ascocarp Anatomy: Episamma  

622. Episamma presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

623. Episamma structure    ()  1. crystalline ()

Ascocarp Anatomy: Epithecium  

627. Epithecium presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

639. Epithecial hyphae apical cell pigmentation {in H2O}     1. hyaline (translucent)     25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine)     27. brownish yellow (= yellowish brown)
(Select a character state: 1 25 27 to add this modifier: )
(Select a character state: 1 25 27 to add this note/comment: )

Ascocarp Anatomy: Hymenium  

659. Hymenium gel presence     1. absent     2. present (interascal hyphae gelatinized)
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

667. Hymenium euamyloid reaction     1. Lugol’s negative     2. Lugol’s positive
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

Ascocarp Anatomy: Interascal Hyphae  

673. Interascal hyphae presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

675. Interascal hyphae orientation     1. irregular     2. anticlinal
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

679. Interascal hyphae cell length [numerical]

Ascocarp Anatomy: Asci  

751. Ascus shape     3. sacculiform     4. broadly clavate     5. clavate     6. narrowly clavate     8. cylindrical
(Select a character state: 3 4 5 6 8 to add this modifier: )
(Select a character state: 3 4 5 6 8 to add this note/comment: )

753. Ascus length [numerical]

769. Ascus dehiscence type    ()  4. verrucarial ()

Ascocarp Anatomy: Ascospores  

783. Ascospore arrangement     1. not uniseriate     2. uniseriate
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

784. Ascospore number [categorical]     1. 1–2 per ascus     2. c. 4 per ascus     3. c. 8 per ascus     4. 12-16 per ascus ()
(Select a character state: 1 2 3 4 to add this modifier: )
(Select a character state: 1 2 3 4 to add this note/comment: )

785. Ascospore number [numerical]

786. Ascospore shape     2. subglobose     3. oval     4. ellipsoid     5. broadly ellipsoid     16. fusiform
(Select a character state: 2 3 4 5 16 to add this modifier: )
(Select a character state: 2 3 4 5 16 to add this note/comment: )

787. Ascospore length [numerical]

792. Ascospore septum presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

793. Ascospore septum orientation     1. transversely septate     2. muriform
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

795. Ascospore {median} septum number [numerical]

802. Ascospore wall width [categorical]     1. thin     2. thick
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

803. Ascospore wall width [numerical]

804. Ascospore wall differentiation {visible in the light microscope}    ()  2. distinctly differentiated (into primary and secondary wall) ()

810. Ascospore wall pigmentation     1. hyaline (translucent)     2. grey     4. pale brown (= brownish white, whitish brown)     5. middle brown     7. dark brown (= brownish black, blackish brown)
(Select a character state: 1 2 4 5 7 to add this modifier: )
(Select a character state: 1 2 4 5 7 to add this note/comment: )

812. Ascospore perispore ornamentation     1. not ornamented (psilate, smooth)     2. ornamented
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

Conidioma Morphology  

816. Conidiomata presence [mandatory character]     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

817. Conidiomata kind    ()  1. pycnidial ()

818. Conidiomata abundance     1. sparse     2. moderately abundant     3. strongly abundant
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

Conidioma Anatomy: Conidia  

873. Conidium shape     10. oblong-obtuse     11. bacilliform
(Select a character state: 10 11 to add this modifier: )
(Select a character state: 10 11 to add this note/comment: )

879. Conidium length [numerical]

Chemistry: Secondary Metabolites  

913. Known secondary metabolites: presence    ()  1. absent ()

Chemistry: Spot Tests  

926. Spot test reactions    ()  1. absent ()