Taxon name:

Data maintenance  

1. Data set source [mandatory] [mandatory character]     1. authentic     2. compiled     3. template     5. standard item
(Select a character state: 1 2 3 5 to add this modifier: )
(Select a character state: 1 2 3 5 to add this note/comment: )

2. Data set author(s) [mandatory] [mandatory character]     6. Bowler P. A.     30. Kashiwadani H.     42. Nash T.H.  None selected () ()

3. Data set reviewer(s)    ()  59. Schramm R. & Nash T.H. ()

4. Data set revision status [mandatory character]     2. revised  None selected () ()

5. Data set publication permission [mandatory character]     1. no  None selected () ()

Nomenclature: Taxonomic Hierarchy  

7. Current taxonomic status     1. accepted     2. basionymous
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

8. Taxonomic rank [mandatory] [mandatory character]     2. species  None selected () ()

11. Genus name [mandatory] [mandatory character]     637. Ramalina  None selected () ()

12. Synonyms

13. Family name [mandatory] [mandatory character]     111. Ramalinaceae Ag. (1821)  None selected () ()

15. Order name     9. Lecanorales  None selected () ()

Nomenclature: Type Information  

17. Type



Nomenclature: Taxonomic Literature  

21. Taxonomic notes


22. Taxonomic literature

Biogeography  

23. Hemisphere distribution    ()  1. Northern hemispheric ()

24. Floristic province {= zonal distribution} [mandatory] [mandatory character]     1. holarctic     2. arctic     3. subarctic     4. boreal     5. Mediterranean     6. subtropical     7. pantropical     8. neotropical     9. palaeotropic     10. subantarctic     11. antarctic     14. cosmopolitan
(Select a character state: 1 2 3 4 5 6 7 8 9 10 11 14 to add this modifier: )
(Select a character state: 1 2 3 4 5 6 7 8 9 10 11 14 to add this note/comment: )

25. Altitude [mandatory] [mandatory character]     1. coastal     2. colline     3. submontane     4. montane     5. upper montane     6. subalpine     7. alpine
(Select a character state: 1 2 3 4 5 6 7 to add this modifier: )
(Select a character state: 1 2 3 4 5 6 7 to add this note/comment: )

38. Checklist records [mandatory] [mandatory character]     15. Mexico     20. Sonoran Desert     26. United States and Canada (continental)
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(Select a character state: 15 20 26 to add this note/comment: )

Ecology  

43. Life habit    ()  2. biotroph (inhabiting living organisms) ()

44. Lifestyle [mandatory] [mandatory character]    ()  5. lichenicolous (= parasitic on lichens) ()

45. Substrate preference [mandatory character]     1. terricolous (on soil)     2. bryophytic (on bryophytes)     3. lignicolous (on wood)     4. corticolous (on bark)     5. epiphyllous (on living leaves)     6. saxicolous (on rock)
(Select a character state: 1 2 3 4 5 6 to add this modifier: )
(Select a character state: 1 2 3 4 5 6 to add this note/comment: )

53. Rock substrate mineral content     1. non-calciferous (lacking calcites)     2. calciferous (containing calcites)
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

Host or Phorophyte Taxonomy  

71. Host fungus or lichen species

Thallus Morphology: General Morphology  

104. Thallus growth form [mandatory character]     2. crustose     3. foliose     4. fruticose
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(Select a character state: 2 3 4 to add this note/comment: )

105. Thallus primary subdivision     1. continuous     2. divided
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

106. Thallus secondary subdivision     1. entire     2. subdivided
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

108. Crustose growth organization     2. granular     8. subfruticose (minutely fruticose, dactyliform)
(Select a character state: 2 8 to add this modifier: )
(Select a character state: 2 8 to add this note/comment: )

109. Crustose growth structure     1. bullate     2. subsquamulose
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

112. Foliose growth organization    ()  1. lobed ()

113. Foliose lobe widening     1. widening     2. parallel
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

116. Foliose lobe orientation     1. horizontal     2. ascending     3. imbricate
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(Select a character state: 1 2 3 to add this note/comment: )

117. Foliose lobe front    ()  1. angular ()

118. Foliose lobe curvature     1. concave     2. subconcave     3. plane     4. subconvex     5. convex
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(Select a character state: 1 2 3 4 5 to add this note/comment: )

119. Fruticose growth organization     1. erect     4. subpendulous     5. pendulous
(Select a character state: 1 4 5 to add this modifier: )
(Select a character state: 1 4 5 to add this note/comment: )

121. Fruticose branch structure     1. terete     2. subterete     3. applanate (moderately flattened)     4. compressed (strongly flattened)     5. alate     6. striate     7. ridged
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(Select a character state: 1 2 3 4 5 6 7 to add this note/comment: )

123. Fruticose branch segmentation     1. not segmented     2. segmented
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

126. Thallus parts imbrication     1. not imbricate     2. imbricate
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

129. Thallus parts length [categorical]     1. short     2. long
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

130. Thallus parts length [numerical]

131. Thallus parts width [categorical]     1. narrow     2. wide
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

132. Thallus parts width [numerical]

133. Thallus parts thickness [categorical]     1. thin     2. thick
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

134. Thallus parts thickness [numerical]

Thallus Morphology: Thallus Size and Differentiation  

151. Overall thallus diameter [categorical]     1. very small     2. small     3. medium-sized     4. large     5. very large
(Select a character state: 1 2 3 4 5 to add this modifier: )
(Select a character state: 1 2 3 4 5 to add this note/comment: )

153. Overall thallus length [numerical]

154. Thallus joints main branch {basal} constriction     1. basally not constricted     2. basally constricted
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

155. Thallus joints secondary branch {basal} constriction     1. basally not constricted     2. basally constricted
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

157. Thallus apex width [numerical]

Thallus Morphology: Outline  

164. Thallus margin structure     1. entire     2. sulcate     3. dentate     4. laciniate     5. cucculate     6. undulate     7. isidiate     8. phyllidiate     9. crenulate     10. dissected     11. lobulate
(Select a character state: 1 2 3 4 5 6 7 8 9 10 11 to add this modifier: )
(Select a character state: 1 2 3 4 5 6 7 8 9 10 11 to add this note/comment: )

165. Thallus margin orientation     1. down-turned     2. levelled     3. up-turned
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

Thallus Morphology: Upper Thallus Surface  

171. Upper surface aspect     1. verrucose     2. verruculose
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

172. Upper surface colour     2. white     3. pale grey (= greyish white, whitish grey)     4. grey     6. black     8. pale red (= reddish white, whitish red, incl. pink, rosé)     14. pale green (= greenish white, whitish green)     15. grey-green (= greyish green, greenish gray)     16. brownish green (= greenish brown, incl. olive, olivaceous)     17. bluish green (= greenish blue; incl. aeruginose, turquoise)     18. dark green (= blackish green, greenish black)     20. pale blue (= whitish blue, bluish white)     24. yellow     25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine)     26. grey-yellow (= greyish yellow, yellowish grey)     27. brownish yellow (= yellowish brown)     28. greenish yellow (= yellowish green, incl. lime green)     29. dark yellow (= yellowish black, blackish yellow)     31. pale brown (= brownish white, whitish brown)     34. orange (= reddish yellow, yellowish red)
(Select a character state: 2 3 4 6 8 14 15 16 17 18 20 24 25 26 27 28 29 31 34 to add this modifier: )
(Select a character state: 2 3 4 6 8 14 15 16 17 18 20 24 25 26 27 28 29 31 34 to add this note/comment: )

173. Upper surface convolution    ()  1. convoluted ()

174. Upper surface curvature     1. plane     2. subconcave     3. concave     4. subconvex     5. convex
(Select a character state: 1 2 3 4 5 to add this modifier: )
(Select a character state: 1 2 3 4 5 to add this note/comment: )

175. Upper surface glossiness     1. matt (opaque, dull)     2. glossy (shiny)     3. vitreous
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

176. Upper surface roughness     1. smooth     2. rough
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

178. Upper surface special structures: presence     1. absent     2. present:
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

179. Upper surface fissures: presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

180. Upper surface fissures: arrangement     1. single     2. reticulate
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

183. Upper surface fissures: location     1. central     2. peripheral
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

184. Upper surface fissures: orientation     1. longitudinal     2. transverse     3. without predominant orientation     4. both transverse and longitudinal
(Select a character state: 1 2 3 4 to add this modifier: )
(Select a character state: 1 2 3 4 to add this note/comment: )

185. Upper surface fissures: position     1. on tubercles     2. on wrinkles     3. on folds     4. within pseudocyphellae     5. between wrinkles
(Select a character state: 1 2 3 4 5 to add this modifier: )
(Select a character state: 1 2 3 4 5 to add this note/comment: )

186. Upper surface pits: presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

189. Upper surface folds: presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

190. Upper surface folds: orientation     1. without particular orientation     2. longitudinally orientated     3. transversally orientated
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

193. Upper surface patches: presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

195. Upper surface patches: outline     1. punctiform     2. orbicular     3. linear
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

196. Upper surface pruina: presence     1. epruinose     2. pruinose
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

197. Upper surface pruina: abundance     1. scarce     2. abundant
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

198. Upper surface pruina: colour     2. greyish     4. greenish     5. yellowish     8. whitish
(Select a character state: 2 4 5 8 to add this modifier: )
(Select a character state: 2 4 5 8 to add this note/comment: )

218. Upper surface cilia: presence     1. eciliate     2. ciliate
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

220. Upper surface cilia: position     2. apical     6. marginal
(Select a character state: 2 6 to add this modifier: )
(Select a character state: 2 6 to add this note/comment: )

222. Upper surface cilia: length [numerical]

224. Upper surface cilia: apical brightness [categorical]     1. bright tipped     2. evenly coloured     3. dark tipped
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

230. Upper surface isidia: presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

234. Upper surface isidia: arrangement     1. isolated     2. clustered
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

235. Upper surface isidia: shape     4. inflated     5. cylindrical     7. spathulate
(Select a character state: 4 5 7 to add this modifier: )
(Select a character state: 4 5 7 to add this note/comment: )

237. Upper surface isidia: ramification     1. not branched     2. branched
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

243. Upper surface papillae: presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

244. Upper surface papillae: shape     1. minute (barely distinct)     2. subhemispherical     3. verrucose     4. cylindrical
(Select a character state: 1 2 3 4 to add this modifier: )
(Select a character state: 1 2 3 4 to add this note/comment: )

252. Upper surface soralia: presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

253. Upper surface soralia: diameter [categorical]     1. minute     2. broad
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

254. Upper surface soralia: diameter [numerical]

255. Upper surface soralia: colour     3. pale grey (= greyish white, whitish grey)     8. pale red (= reddish white, whitish red, incl. pink, rosé)     14. pale green (= greenish white, whitish green)     16. brownish green (= greenish brown, incl. olive, olivaceous)     20. pale blue (= whitish blue, bluish white)     25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine)     26. grey-yellow (= greyish yellow, yellowish grey)     27. brownish yellow (= yellowish brown)     28. greenish yellow (= yellowish green, incl. lime green)     29. dark yellow (= yellowish black, blackish yellow)     31. pale brown (= brownish white, whitish brown)     34. orange (= reddish yellow, yellowish red)
(Select a character state: 3 8 14 16 20 25 26 27 28 29 31 34 to add this modifier: )
(Select a character state: 3 8 14 16 20 25 26 27 28 29 31 34 to add this note/comment: )

256. Upper surface soralia: location     1. central     2. laminal     3. subperipheral     4. peripheral
(Select a character state: 1 2 3 4 to add this modifier: )
(Select a character state: 1 2 3 4 to add this note/comment: )

257. Upper surface soralia: position     1. apical     2. subapical     3. lateral
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

258. Upper surface soralia: origin     1. "ab initio" (literally: from the the beginning, i.e., directly forming on anywhere on thallus surface, breaking through the cortex)     2. papillar (forming at the tips of papillae, i.e., elongated, finger-like projections)     3. tubercular (eroding from the tip of tubercles, warts)     4. pustular (erupting from pustules)     5. nodular (developing on contorted, knot-like structures of branches of fruticose lichens)     6. rugular (originating on wrinkles)     7. on folds (forming on thallus folds)     8. fissural (forming from cracks or fissures)     9. undular (forming on thallus undulations)     10. along scars of fibrils
(Select a character state: 1 2 3 4 5 6 7 8 9 10 to add this modifier: )
(Select a character state: 1 2 3 4 5 6 7 8 9 10 to add this note/comment: )

259. Upper surface soralia: shape     1. semicapitate     2. capitate     3. hooded (helmet-shaped)     4. spathulate     6. maculiform
(Select a character state: 1 2 3 4 6 to add this modifier: )
(Select a character state: 1 2 3 4 6 to add this note/comment: )

260. Upper surface soralia: outline     1. erose (diffuse, unspecific)     2. punctiform     3. irregular ("effigurate")     4. linear
(Select a character state: 1 2 3 4 to add this modifier: )
(Select a character state: 1 2 3 4 to add this note/comment: )

261. Upper surface soralia: elevation     1. concave     2. plane     3. prominent
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

262. Upper surface soralia: excavation    ()  2. fully excavated ()

265. Upper surface soralia: arrangement     1. in isolated lines     2. reticulate fused
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

266. Upper surface soralia: orientation     1. longitudinal     2. transversal
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

268. Upper surface soralia: development {from isidia}     1. not isidiate     2. isidiate
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

271. Upper surface tubercles: presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

272. Upper surface tubercles: size [categorical]     1. very small     2. small     3. large
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

Thallus Anatomy: Upper Cortex  

329. Upper cortex: presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

330. Upper cortex type     1. phenocortical     2. eucortical
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

332. Upper cortex height [numerical]

334. Upper cortex plectenchyma structure     1. prosoplectenchymatous     2. paraplectenchymatous
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

335. Upper cortex paraplectenchyma type    ()  1. textura globularis ()

337. Upper cortex hyphal orientation    ()  2. anticlinal ()

338. Upper cortex layer layer differentiation     1. one     2. two
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

339. Upper cortex hyphal cell structure    ()  1. chondroid ()

342. Upper cortex hyphal cell strands     1. conical hyphal cell strands penetrating the photobiont layer     2. irregular hyphal cell strands     3. without distinct strands of hyphae
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

349. Upper cortex pigment location     1. in upper layer     2. in lower layer
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

360. Upper cortex epicortex: presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

367. Upper cortex outer layer: structure     1. prosoplectenchymatous     2. paraplectenchymatous     3. scleroplectenchymatous
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

371. Upper cortex inner layer: structure     1. prosoplectenchymatous     2. paraplectenchymatous     3. scleroplectenchymatous
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

Thallus Anatomy: Photobiont Layer  

374. Photobiont layer: height [numerical]

376. Photobiont layer: compartmentalization     1. continuous     2. clustered     3. in vaulted chambers     4. scattered
(Select a character state: 1 2 3 4 to add this modifier: )
(Select a character state: 1 2 3 4 to add this note/comment: )

Thallus Anatomy: Medulla  

380. Medulla height [numerical]

384. Medulla density     1. compact     2. dense     3. loose     4. very loose, becoming hollow
(Select a character state: 1 2 3 4 to add this modifier: )
(Select a character state: 1 2 3 4 to add this note/comment: )

388. Medulla hyphal arrangement     1. forming several bundles     2. forming a compact mass
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

389. Medulla hyphal orientation     1. predominantly along the lobe axis     2. without particular orientation in upper part, predominantly along the lobe axis in lower part     3. without particular orientation
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

393. Medulla euamyloid reactivity     1. in Lugol's solution negative     2. in Lugol's solution positive
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

395. Medulla iodine reaction intensity     1. weakly euamyloid     2. strongly euamyloid
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

396. Medulla iodine reaction colour     1. I + blue     2. I + violet
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

Reproduction  

414. Ascomata presence [mandatory] [mandatory character]     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

Ascocarp Morphology: General Ascocarp Morphology  

421. Ascocarp location {on main thallus}     1. central     2. laminal     3. subperipheral     4. peripheral
(Select a character state: 1 2 3 4 to add this modifier: )
(Select a character state: 1 2 3 4 to add this note/comment: )

424. Ascocarp position {on primary thallus parts}     1. central     2. laminal     3. subperipheral     4. peripheral     5. subterminal     6. terminal
(Select a character state: 1 2 3 4 5 6 to add this modifier: )
(Select a character state: 1 2 3 4 5 6 to add this note/comment: )

429. Ascocarp attachment     1. adnate (semi-immersed)     2. sessile     3. substipitate     4. stipitate
(Select a character state: 1 2 3 4 to add this modifier: )
(Select a character state: 1 2 3 4 to add this note/comment: )

430. Ascocarp basis     1. not constricted (flat below)     2. slightly constricted (moderately conical at the base)     3. strongly constricted (strongly conical at the base)
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

431. Ascocarp abundance [categorical]     1. sparse     2. moderately abundant     3. abundant
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

437. Ascocarp size [categorical]     1. small     2. large
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

438. {orbicular} ascocarp diameter [numerical]

Ascocarp Morphology: Margin  

448. Ascocarp margin: structure     2. lobate     3. lobulate     4. crenulate     5. granulose     7. verrucose     8. verruculose
(Select a character state: 2 3 4 5 7 8 to add this modifier: )
(Select a character state: 2 3 4 5 7 8 to add this note/comment: )

449. Ascocarp margin: roughness     1. smooth     5. rough
(Select a character state: 1 5 to add this modifier: )
(Select a character state: 1 5 to add this note/comment: )

450. Ascocarp margin: outline     1. not flexuose     2. slightly flexuose     3. strongly flexuose
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

452. Ascocarp margin: development     1. indistinct (thin)     3. prominent (thick)
(Select a character state: 1 3 to add this modifier: )
(Select a character state: 1 3 to add this note/comment: )

453. Ascocarp margin: colour     3. pale grey (= greyish white, whitish grey)     5. dark gray (= greyish black, blackish grey)     8. pale red (= reddish white, whitish red, incl. pink, rosé)     9. grey-red (= greyish red, reddish grey)     14. pale green (= greenish white, whitish green)     15. grey-green (= greyish green, greenish gray)     16. brownish green (= greenish brown, incl. olive, olivaceous)     17. bluish green (= greenish blue; incl. turquoise, aeruginose)     18. dark green (= blackish green, greenish black)     20. pale blue (= whitish blue, bluish white)     21. grey-blue (= bluish grey, greyish blue)     25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine)     26. grey-yellow (= greyish yellow, yellowish grey)     27. brownish yellow (= yellowish brown)     28. greenish yellow (= yellowish green, incl. lime green)     31. pale brown (= brownish white, whitish brown)     32. grey-brown (= greyish brown, brownish grey)     34. orange (= reddish yellow, yellowish red)
(Select a character state: 3 5 8 9 14 15 16 17 18 20 21 25 26 27 28 31 32 34 to add this modifier: )
(Select a character state: 3 5 8 9 14 15 16 17 18 20 21 25 26 27 28 31 32 34 to add this note/comment: )

454. Ascocarp margin: pigment distribution     1. brighter than thallus     2. concolourous with thallus     3. darker than thallus     4. brighter than disk     5. concolourous with disk     6. darker than disk
(Select a character state: 1 2 3 4 5 6 to add this modifier: )
(Select a character state: 1 2 3 4 5 6 to add this note/comment: )

457. Ascocarp margin: width [numerical]

458. Ascocarp margin: pruina abundance     1. epruinose     2. slightly pruinose     3. distinctly pruinose
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

462. Ascocarp margin: surface structures presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

463. Ascocarp margin: soredia     1. absent     2. present (sorediate)
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

464. Ascocarp margin: isidia     1. absent     2. present (isidiate)
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

466. Ascocarp margin: cilia     1. absent     2. present (ciliate)
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

Ascocarp Morphology: Disk  

487. Ascocarp disk: curvature     1. urceolate (strongly concave)     2. excavate (concave)     3. plane (expanded)     4. weakly convex     5. strongly convex
(Select a character state: 1 2 3 4 5 to add this modifier: )
(Select a character state: 1 2 3 4 5 to add this note/comment: )

488. Ascocarp disk: colour     3. pale grey (= greyish white, whitish grey)     5. dark gray (= greyish black, blackish grey)     8. pale red (= reddish white, whitish red, incl. pink, rosé)     9. grey-red (= greyish red, reddish grey)     14. pale green (= greenish white, whitish green)     15. grey-green (= greyish green, greenish gray)     16. brownish green (= greenish brown, incl. olive, olivaceous)     17. bluish green (= greenish blue; incl. aeruginose, turquoise)     18. dark green (= blackish green, greenish black)     20. pale blue (= whitish blue, bluish white)     21. grey-blue (= bluish grey, greyish blue)     25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine)     26. grey-yellow (= greyish yellow, yellowish grey)     27. brownish yellow (= yellowish brown)     28. greenish yellow (= yellowish green, incl. lime green)     29. dark yellow (= yellowish black, blackish yellow)     31. pale brown (= brownish white, whitish brown)     32. grey-brown (= greyish brown, brownish grey)     34. orange (= reddish yellow, yellowish red)
(Select a character state: 3 5 8 9 14 15 16 17 18 20 21 25 26 27 28 29 31 32 34 to add this modifier: )
(Select a character state: 3 5 8 9 14 15 16 17 18 20 21 25 26 27 28 29 31 32 34 to add this note/comment: )

491. Ascocarp disk: pruina presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

492. Ascocarp disk: pruina abundance     1. scarce     2. abundant
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

493. Ascocarp disk: pruina colour     1. white     2. yellow greenish     3. green     4. yellow     5. red     6. brownish red (= reddish brown)     7. pale grey (= whitish grey, greyish white)
(Select a character state: 1 2 3 4 5 6 7 to add this modifier: )
(Select a character state: 1 2 3 4 5 6 7 to add this note/comment: )

494. Ascocarp umbo presence    ()  1. absent ()

495. Ascocarp umbo abundance     1. occasional     2. regular
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

Ascocarp Anatomy: Episamma  

622. Episamma presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

Ascocarp Anatomy: Epithecium  

639. Epithecial hyphae apical cell pigmentation {in H2O}     3. pale grey (= whitish grey, greyish white)     8. pale red (= reddish white, whitish red, incl. pink, rosé)     10. brownish red (= reddish brown)     14. pale green (= greenish white, whitish green)     16. brownish green (= greenish brown, incl. olive, olivaceous)     20. pale blue (= whitish blue, bluish white)     22. brownish blue (= bluish brown)     25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine)     26. grey-yellow (= greyish yellow, yellowish grey)     27. brownish yellow (= yellowish brown)     28. greenish yellow (= yellowish green, incl. lime green)     29. dark yellow (= blackish yellow, yellowish black)     31. pale brown (= brownish white, whitish brown)     32. grey-brown (= greyish brown, brownish grey)     33. dark brown (= brownish black, blackish brown)     34. orange (= reddish yellow, yellowish red)
(Select a character state: 3 8 10 14 16 20 22 25 26 27 28 29 31 32 33 34 to add this modifier: )
(Select a character state: 3 8 10 14 16 20 22 25 26 27 28 29 31 32 33 34 to add this note/comment: )

Ascocarp Anatomy: Hymenium  

653. Hymenium height [numerical]

Ascocarp Anatomy: Interascal Hyphae  

676. Interascal hyphae ramification     1. scarcely branched     2. distinctly branched
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

677. Interascal hyphae anastomoses     1. not or scarcely anastomosed     2. distinctly anastomosed
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

Ascocarp Anatomy: Asci  

751. Ascus shape     1. globose     4. broadly clavate     5. clavate     6. narrowly clavate     7. obclavate     8. cylindrical
(Select a character state: 1 4 5 6 7 8 to add this modifier: )
(Select a character state: 1 4 5 6 7 8 to add this note/comment: )

753. Ascus length [numerical]

Ascocarp Anatomy: Ascospores  

783. Ascospore arrangement     1. not uniseriate     2. uniseriate
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

786. Ascospore shape     1. globose     2. subglobose     3. oval     4. ellipsoid     5. broadly ellipsoid     6. ovoid     7. obovoid     10. oblong-obtuse     13. cylindrical     16. fusiform     28. semilunate     31. reniform (incl. fabiform)     32. curved     35. allantoid     37. sigmoid
(Select a character state: 1 2 3 4 5 6 7 10 13 16 28 31 32 35 37 to add this modifier: )
(Select a character state: 1 2 3 4 5 6 7 10 13 16 28 31 32 35 37 to add this note/comment: )

787. Ascospore length [numerical]

790. Ascospore attenuation     1. obtuse     2. aciculate
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

803. Ascospore wall width [numerical]

Conidioma Morphology  

816. Conidiomata presence [mandatory character]     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

820. Conidiomata length [numerical]

824. Conidiomata position {on thallus parts}     1. on tubercles (warts)     2. on wrinkles     3. on folds     4. on simple isidia     5. on branched isidia     6. on projections     7. on muri     8. on primary squamules     9. on podetia     10. on margins of scyphi     11. on margins of ascocarps     12. in primordia of ascocarps     13. in the hymenium of ascocarps
(Select a character state: 1 2 3 4 5 6 7 8 9 10 11 12 13 to add this modifier: )
(Select a character state: 1 2 3 4 5 6 7 8 9 10 11 12 13 to add this note/comment: )

825. Conidiomata general position     1. lateral (along the margin of thallus parts)     2. subapical (just below the tips of thallus parts)     3. apical
(Select a character state: 1 2 3 to add this modifier: )
(Select a character state: 1 2 3 to add this note/comment: )

826. Conidiomata location     1. central (on thallus)     2. laminal (on thallus)     3. subperipheral (on thallus)     4. peripheral (on thallus)
(Select a character state: 1 2 3 4 to add this modifier: )
(Select a character state: 1 2 3 4 to add this note/comment: )

827. Conidiomata supporting structure: presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

Conidioma Anatomy: Pycnidia  

833. Pycnidium shape     1. globular     2. subglobular     4. ovoid     6. pyriform     7. ampulliform     11. bowl-like     15. cerebriform
(Select a character state: 1 2 4 6 7 11 15 to add this modifier: )
(Select a character state: 1 2 4 6 7 11 15 to add this note/comment: )

835. Pycnidium compartmentalization     1. unilocular     2. plurilocular
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

Conidioma Anatomy: Conidia  

873. Conidium shape     1. globose     2. subglobose     3. oval     4. ellipsoid     5. broadly ellipsoid     13. cylindrical     16. fusiform     17. filiform     37. bifusiform
(Select a character state: 1 2 3 4 5 13 16 17 37 to add this modifier: )
(Select a character state: 1 2 3 4 5 13 16 17 37 to add this note/comment: )

879. Conidium length [numerical]

Chemistry: Secondary Metabolites  

913. Known secondary metabolites: presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

914. Known secondary metabolites: location     1. throughout the cortex     2. only in the upper part of the cortex     3. only in the lower part of the cortex     4. throughout the thallus medulla     5. only in the upper medulla     6. only in the lower medulla     7. in the ascocarp margin     8. within the subhymenium or hypothecium     9. throughout the whole central axis     10. restricted to the outer part of the central axis     11. in soredia (or soralia)
(Select a character state: 1 2 3 4 5 6 7 8 9 10 11 to add this modifier: )
(Select a character state: 1 2 3 4 5 6 7 8 9 10 11 to add this note/comment: )

915. Known secondary metabolites: name     133. 4-O-demethylbarbatic acid     235. barbatic acid     240. bourgeanic acid     267. consalazinic acid     289. divaricatic acid     311. evernic acid     356. hypoprotocetraric acid     414. methyl 3,5-dichlorolecanorate     458. nordivaricatic acid     470. norstictic acid     512. protocetraric acid     536. salazinic acid     546. sekikaic acid     549. skyrin     559. stictic acid     586. usnic acid
(Select a character state: 133 235 240 267 289 311 356 414 458 470 512 536 546 549 559 586 to add this modifier: )
(Select a character state: 133 235 240 267 289 311 356 414 458 470 512 536 546 549 559 586 to add this note/comment: )

917. Unknown secondary metabolites: presence     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

Chemistry: Spot Tests  

926. Spot test reactions     1. absent     2. present
(Select a character state: 1 2 to add this modifier: )
(Select a character state: 1 2 to add this note/comment: )

927. Upper surface K reactivity     1. – (negative)     2. + deep yellow     3. + bright yellow (citrine, lemon)     4. + yellow changing to red     5. + red     6. + blackish red     7. + bluish red     8. + orange     9. + pink
(Select a character state: 1 2 3 4 5 6 7 8 9 to add this modifier: )
(Select a character state: 1 2 3 4 5 6 7 8 9 to add this note/comment: )

928. Upper surface C reactivity     1. –     2. + deep yellow     3. + bright yellow     4. + yellow changing to red     5. + red     6. + blackish red     7. + bluish red     8. + orange     9. + pink     10. + green
(Select a character state: 1 2 3 4 5 6 7 8 9 10 to add this modifier: )
(Select a character state: 1 2 3 4 5 6 7 8 9 10 to add this note/comment: )

930. Upper surface KC reactivity     1. –     2. + deep yellow     3. + bright yellow (citrine, lemon)     4. + yellow changing to red     5. + red     6. + blackish red     7. + bluish red     8. + orange     9. + pink     10. + green
(Select a character state: 1 2 3 4 5 6 7 8 9 10 to add this modifier: )
(Select a character state: 1 2 3 4 5 6 7 8 9 10 to add this note/comment: )

931. Upper surface PD reactivity     1. –     2. + yellow     3. + bright yellow (citrine, lemon)     4. + yellow changing to red     5. + red     6. + blackish red     7. + bluish red     8. + orange     9. + pink
(Select a character state: 1 2 3 4 5 6 7 8 9 to add this modifier: )
(Select a character state: 1 2 3 4 5 6 7 8 9 to add this note/comment: )

937. Medulla K reactivity     1. –     2. + yellow     3. + bright yellow (citrine, lemon)     4. + yellow changing to red     5. + red     6. + blackish red     7. + bluish red     8. + orange     9. + pink
(Select a character state: 1 2 3 4 5 6 7 8 9 to add this modifier: )
(Select a character state: 1 2 3 4 5 6 7 8 9 to add this note/comment: )

938. Medulla C reactivity     2. + deep yellow     3. + bright yellow (citrine, lemon)     4. + yellow changing to red     5. + red     6. + blackish red     7. + bluish red     8. + orange     9. + pink
(Select a character state: 2 3 4 5 6 7 8 9 to add this modifier: )
(Select a character state: 2 3 4 5 6 7 8 9 to add this note/comment: )

940. Medulla KC reactivity     1. –     2. + yellow     3. + bright yellow (citrine, lemon)     4. + yellow changing to red     5. + red     6. + blackish red     7. + bluish red     8. + pink     9. + orange
(Select a character state: 1 2 3 4 5 6 7 8 9 to add this modifier: )
(Select a character state: 1 2 3 4 5 6 7 8 9 to add this note/comment: )

941. Medulla PD reactivity     1. –     2. + deep yellow     3. + bright yellow (citrine, lemon)     4. + yellow changing to red     5. + red     6. + blackish red     7. + bluish red     8. + pink     9. + orange
(Select a character state: 1 2 3 4 5 6 7 8 9 to add this modifier: )
(Select a character state: 1 2 3 4 5 6 7 8 9 to add this note/comment: )

947. Soredia K reactivity     1. –     2. + deep yellow     3. + pink     4. + orange
(Select a character state: 1 2 3 4 to add this modifier: )
(Select a character state: 1 2 3 4 to add this note/comment: )