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Go to: Data maintenance    Nomenclature: Taxonomic Hierarchy    Nomenclature: Type Information    Nomenclature: Taxonomic Literature    Biogeography    Ecology    Lichen Photobionts    Thallus Morphology: General Morphology    Thallus Morphology: Thallus Size and Differentiation    Thallus Morphology: Outline    Thallus Morphology: Upper Thallus Surface    Thallus Morphology: Lower Thallus Surface    Thallus Anatomy: Upper Cortex    Reproduction    Ascocarp Morphology: General Ascocarp Morphology    Ascocarp Morphology: Margin    Ascocarp Morphology: Disk    Ascocarp Anatomy: Exciple    Ascocarp Anatomy: Epithecium    Ascocarp Anatomy: Hymenium    Ascocarp Anatomy: Subhymenium    Ascocarp Anatomy: Hypothecium    Ascocarp Anatomy: Asci    Ascocarp Anatomy: Ascospores    Conidioma Morphology    Conidioma Anatomy: Conidiogeneous Cells    Conidioma Anatomy: Conidia    Chemistry: Secondary Metabolites    Chemistry: Spot Tests    Index of all items of the project


Taxon name:



Data maintenance   back to top

1. Data set source [mandatory] [mandatory character]     2. compiled     3. template     5. standard item
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2. Data set author(s) [mandatory] [mandatory character]     42. Nash T.H.     52. Schramm R. & Nash T. H.  None selected () ()
3. Data set reviewer(s) [mandatory character]     45. Nash T.H.     59. Schramm R. & Nash T.H.
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4. Data set revision status [mandatory character]     1. not revised     2. revised  None selected () ()
5. Data set publication permission [mandatory character]     1. no     2. yes  None selected () ()
6. Data set publication date



Nomenclature: Taxonomic Hierarchy   back to top

7. Current taxonomic status     1. accepted     2. basionymous
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8. Taxonomic rank [mandatory] [mandatory character]     2. species  None selected () ()
11. Genus name [mandatory] [mandatory character]     343. Koerberia     494. Pannaria  None selected () ()
12. Synonyms


13. Family name [mandatory] [mandatory character]     90. Pannariaceae Tuck. (1872)  None selected () ()
14. Suborder name     6. Peltigerineae  None selected () ()
15. Order name     9. Lecanorales  None selected () ()

Nomenclature: Type Information   back to top

17. Type [mandatory character]


19. Protologue


20. Type specimen label data



Nomenclature: Taxonomic Literature   back to top

21. Taxonomic notes


22. Taxonomic literature



Biogeography   back to top

23. Hemisphere distribution    ()  1. Northern hemispheric ()
24. Floristic province {= zonal distribution} [mandatory] [mandatory character]     1. holarctic     2. arctic     3. subarctic     4. boreal     5. Mediterranean     6. subtropical     7. pantropical     8. neotropical     9. palaeotropic     10. subantarctic     11. antarctic     14. cosmopolitan
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25. Altitude [mandatory] [mandatory character]     1. coastal     2. colline     3. submontane     4. montane     5. upper montane     6. subalpine     7. alpine
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27. Continent distribution [mandatory] [mandatory character]     1. Africa     2. Antarctica     3. Asia-Temperate     4. Australasia     5. Europe     6. Northern America     7. Southern America     8. Pacific     9. Asia-Tropical
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28. Major geographic region     3. South-western Europe     6. Northern Africa     41. South-western U.S.A.
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29. European distribution records     19. France (excl. Corsica)     20. Portugal     26. Italy (incl. San Marino & The Vatican City, excl. Sicily, Sardinia)
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30. African distribution records    ()  4. Morocco ()
35. North American distribution records     46. Arizona (USA)     47. California (USA)
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38. Checklist records [mandatory]     13. Italy     15. Mexico     16. Morocco     20. Sonoran Desert     26. United States and Canada (continental)
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Ecology   back to top

43. Life habit    ()  2. biotroph (inhabiting living organisms) ()
44. Lifestyle [mandatory] [mandatory character]    ()  4. lichenized (= parasitic and / or mutual symbiotic with photobionts) ()
45. Substrate preference [mandatory character]     1. terricolous (on soil)     2. bryophytic (on bryophytes)     3. lignicolous (on wood)     4. corticolous (on bark)     6. saxicolous (on rock)
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47. Habitat insolation     1. sun-exposed     2. partially shaded     3. fully shaded
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48. Habitat moisture     1. dry     2. moist     3. wet
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53. Rock substrate mineral content     1. non-calciferous (lacking calcites)     2. calciferous (containing calcites)
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Lichen Photobionts   back to top

86. Primary photobiont genus     6. Calothrix     32. Myrmecia     35. Nostoc     42. Scytonema     43. Stichococcus     44. Stigonema
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91. Secondary photobiont presence     1. absent     2. present
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98. Secondary photobiont genus    ()  35. Nostoc ()

Thallus Morphology: General Morphology   back to top

104. Thallus growth form [mandatory character]     1. indistinct     2. crustose     3. foliose
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105. Thallus primary subdivision     1. continuous     2. divided
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107. Thallus ramification pattern     4. subdichotomous     5. irregular
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108. Crustose growth organization     1. leprose     2. granular     3. rimose     5. areolate (primarily areolate)     6. placodioid     7. squamulose
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109. Crustose growth structure     1. bullate     2. subsquamulose
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110. Crustose growth aggregation    ()  2. scattered (dispersed) ()
111. Crustose growth continuity     1. continuous throughout     2. dispersed along the periphery, continuous in the centre     3. dispersed throughout and distinctly seperate
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112. Foliose growth organization    ()  1. lobed ()
114. Foliose lobe outline     1. elongate     3. linear
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115. Foliose lobe separation     1. not laciniate     2. laciniate
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116. Foliose lobe orientation     1. horizontal     2. ascending     3. imbricate
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117. Foliose lobe front     1. angular     2. rounded
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121. Fruticose branch structure     1. terete     2. subterete     3. applanate (moderately flattened)     4. compressed (strongly flattened)     6. striate     7. ridged
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122. Fruticose branch flattening     1. bifacial     2. unifacial
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126. Thallus parts imbrication     1. not imbricate     2. imbricate
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128. Thallus parts distance [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
mm

130. Thallus parts length [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
mm long

132. Thallus parts width [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
mm wide

133. Thallus parts thickness [categorical]     1. thin     2. thick
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134. Thallus parts thickness [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
mm thick


Thallus Morphology: Thallus Size and Differentiation   back to top

151. Overall thallus diameter [categorical]     1. very small     2. small     3. medium-sized     4. large
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152. Overall thallus diameter [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
cm in diameter

157. Thallus apex width [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
mm wide

159. Thallus apex outline     1. rounded     2. truncate
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Thallus Morphology: Outline   back to top

160. Thallus perimeter outline     1. irregular     2. orbicular
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161. Thallus perimeter zonation     1. simple     2. zonate     3. lobed     4. arachnoid     5. fimbriate
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162. Thallus perimeter delimitation     1. indistinct     2. distinct
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164. Thallus margin structure     1. entire     3. dentate     4. laciniate     7. isidiate     8. phyllidiate     9. crenulate     10. dissected     11. lobulate     13. crenate
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167. Primal thallus differentiation     1. peripheral (forming a prothallus)     2. extending (forming a hypothallus)
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168. Primal thallus persistence     1. disappearing     2. persistent
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169. Primal thallus development     1. vestigial     2. pronounced     3. cushion forming
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170. Primal thallus colour     1. black     2. grey     4. green     6. brown     8. white
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Thallus Morphology: Upper Thallus Surface   back to top

171. Upper surface aspect     1. verrucose     2. verruculose
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172. Upper surface colour     4. grey     5. dark gray (= greyish black, blackish grey)     15. grey-green (= greyish green, greenish gray)     16. brownish green (= greenish brown, incl. olive, olivaceous)     17. bluish green (= greenish blue; incl. aeruginose, turquoise)     18. dark green (= blackish green, greenish black)     21. grey-blue (= bluish grey, greyish blue)     23. dark blue (= blackish blue, bluish black)     28. greenish yellow (= yellowish green, incl. lime green)
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174. Upper surface curvature     1. plane     2. subconcave     3. concave     4. subconvex     5. convex
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175. Upper surface glossiness     1. matt (opaque, dull)     2. glossy (shiny)     3. vitreous
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176. Upper surface roughness    ()  1. smooth ()
178. Upper surface special structures: presence     1. absent     2. present:
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196. Upper surface pruina: presence     1. epruinose     2. pruinose
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197. Upper surface pruina: abundance     1. scarce     2. abundant
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198. Upper surface pruina: colour     2. greyish     8. whitish
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230. Upper surface isidia: presence     1. absent     2. present
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231. Upper surface isidia: location    ()  2. laminal ()
232. Upper surface isidia: position {across the surface}     1. marginal     2. submarginal     3. apical
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235. Upper surface isidia: shape    ()  5. cylindrical ()
238. Upper surface isidia: ramification pattern     1. sparse     2. coralloid     3. moderate
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252. Upper surface soralia: presence     1. absent     2. present
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255. Upper surface soralia: colour     2. white     4. grey     30. brown
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256. Upper surface soralia: location     1. central     2. laminal     3. subperipheral     4. peripheral
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257. Upper surface soralia: position    ()  1. apical ()
259. Upper surface soralia: shape     1. semicapitate     2. capitate
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260. Upper surface soralia: outline    ()  1. erose (diffuse, unspecific) ()
262. Upper surface soralia: excavation    ()  2. fully excavated ()
264. Upper surface soralia: differentiation     1. not confluent     2. confluent (but not fusing)
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268. Upper surface soralia: development {from isidia}    ()  2. isidiate ()
269. Upper surface soredia: diameter [categorical]     1. grainy (coarse)     2. farinose (fine)
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275. Upper surface cephalodia: presence     1. absent     2. present
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278. Upper surface lobules: presence     1. absent     2. present
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Thallus Morphology: Lower Thallus Surface   back to top

282. Lower surface overall colour     2. white     4. grey     6. black     30. brown
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286. Lower surface attachment kind     1. by the whole lower surface     2. by holdfasts
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300. Lower surface rhizines: presence     1. absent     2. present (rhizinate)
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302. Lower surface rhizines: abundance     1. sparse     2. moderately abundant     3. abundant
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307. Lower surface rhizines: colour     4. grey     6. black     30. brown
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311. Lower surface rhizines: ramification pattern     1. irregular     2. dichotomous     3. squarrose     4. sparsely furcate
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326. Lower surface tomentum: presence     1. absent     2. present
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Thallus Anatomy: Upper Cortex   back to top

329. Upper cortex: presence     1. absent     2. present
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332. Upper cortex height [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm high

367. Upper cortex outer layer: structure     1. prosoplectenchymatous     2. paraplectenchymatous
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371. Upper cortex inner layer: structure     1. prosoplectenchymatous     2. paraplectenchymatous     3. scleroplectenchymatous
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Reproduction   back to top

414. Ascomata presence [mandatory] [mandatory character]     1. absent     2. present
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Ascocarp Morphology: General Ascocarp Morphology   back to top

421. Ascocarp location {on main thallus}     1. central     2. laminal     3. subperipheral     4. peripheral
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423. Ascocarp position {on primal thallus: pro- or hypothallus}     1. laminal     2. peripheral
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424. Ascocarp position {on primary thallus parts}     5. subterminal     6. terminal
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428. Ascocarp emergence    ()  1. not emerging (immersed, e.g., cryptolecanorine, cryptolecideine, aspicilioid) ()
429. Ascocarp attachment     1. adnate (semi-immersed)     2. sessile     3. substipitate
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431. Ascocarp abundance [categorical]     1. sparse     2. moderately abundant     3. abundant
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437. Ascocarp size [categorical]     1. small     2. large
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438. {orbicular} ascocarp diameter [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
mm in diam.


Ascocarp Morphology: Margin   back to top

447. Ascocarp margin: type     1. lecideine/biatorine     2. lecanorine (incl. zeorine)
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448. Ascocarp margin: structure     2. lobate     3. lobulate     4. crenulate
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449. Ascocarp margin: roughness    ()  1. smooth ()
451. Ascocarp margin: persistency     1. persistent     2. excluded
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452. Ascocarp margin: development     1. indistinct (thin)     2. distinct (moderately thickened)     3. prominent (thick)
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453. Ascocarp margin: colour     4. grey     6. black     13. green     30. brown
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463. Ascocarp margin: soredia     1. absent     2. present (sorediate)
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464. Ascocarp margin: isidia    ()  2. present (isidiate) ()

Ascocarp Morphology: Disk   back to top

488. Ascocarp disk: colour     6. black     7. red     13. green     24. yellow     30. brown
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491. Ascocarp disk: pruina presence     1. absent     2. present
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492. Ascocarp disk: pruina abundance     1. scarce     2. abundant
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493. Ascocarp disk: pruina colour     1. white     4. yellow
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Ascocarp Anatomy: Exciple   back to top

504. Ascocarp excipulum: pigment carbonization     1. biatorine (not carbonized)     2. lecideine (carbonized)
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Ascocarp Anatomy: Epithecium   back to top

627. Epithecium presence     1. absent     2. present
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629. Epithecium height [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm high

639. Epithecial hyphae apical cell pigmentation {in H2O}     10. brownish red (= reddish brown)     16. brownish green (= greenish brown, incl. olive, olivaceous)     17. bluish green (= greenish blue; incl. aeruginose, turquoise)     21. grey-blue (= bluish grey, greyish blue)     23. dark blue (= blackish blue, bluish black)     27. brownish yellow (= yellowish brown)     31. pale brown (= brownish white, whitish brown)
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641. Epithecial hyphae apical cell: pigment solubility {in 10% KOH}     1. insoluble in 10% KOH     2. soluble in 10% KOH
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642. Epithecial hyphae apical cell: pigment reactivity    ()  2. 10% KOH + ()

Ascocarp Anatomy: Hymenium   back to top

653. Hymenium height [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm high

654. Hymenium pigmentation {in H2O}     10. brownish red (= reddish brown)     27. brownish yellow (= yellowish brown)     31. pale brown (= brownish white, whitish brown)
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Ascocarp Anatomy: Subhymenium   back to top

684. Subhymenium height [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm high

699. Subhymenium pigmentation {in H2O}     2. white     4. grey     24. yellow     27. brownish yellow (= yellowish brown)     30. brown
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Ascocarp Anatomy: Hypothecium   back to top

714. Hypothecium plectenchyma structure     1. prosoplectenchymatous     2. paraplectenchymatous
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Ascocarp Anatomy: Asci   back to top

751. Ascus shape     1. globose     4. broadly clavate     5. clavate     6. narrowly clavate     7. obclavate     8. cylindrical
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753. Ascus length [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm long

754. Ascus width [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm wide

758. Ascus tholus iodine reactivity    ()  1. Lugol’s negative ()
760. Ascus tholus iodine reaction colour     1. + red     2. + light blue     3. + dark blue
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761. Ascus tholus amyloidy pattern     1. with amyloid tube     2. with amyloid cap     3. entirely amyloid throughout     5. with non-amyloid narrow axial body and amyloid convergent flanks towards apex
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Ascocarp Anatomy: Ascospores   back to top

784. Ascospore number [categorical]     2. c. 4 per ascus     3. c. 8 per ascus     4. 12-16 per ascus
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786. Ascospore shape     2. subglobose     3. oval     4. ellipsoid     5. broadly ellipsoid     16. fusiform
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787. Ascospore length [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm long

788. Ascospore width [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm wide

792. Ascospore septum presence     1. absent     2. present
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793. Ascospore septum orientation    ()  1. transversely septate ()
795. Ascospore {median} septum number [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
-transversally septate


Conidioma Morphology   back to top

816. Conidiomata presence [mandatory character]     1. absent     2. present
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817. Conidiomata kind    ()  1. pycnidial ()
818. Conidiomata abundance     1. sparse     2. moderately abundant     3. strongly abundant
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Conidioma Anatomy: Conidiogeneous Cells   back to top

865. Conidiogenous cells: position     1. apical     2. intercalar
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Conidioma Anatomy: Conidia   back to top

873. Conidium shape     11. bacilliform     13. cylindrical
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879. Conidium length [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm long

880. Conidium width [numerical]
Min: Lower range: Mean: Upper range: Max: Unit:
µm wide


Chemistry: Secondary Metabolites   back to top

913. Known secondary metabolites: presence     1. absent     2. present
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915. Known secondary metabolites: name     231. atranorin     489. pannaric acid     491. pannarin
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916. Known secondary metabolites: substance class     2. (higher) aliphatic acids     14. (tri-)terpenoids
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917. Unknown secondary metabolites: presence     1. absent     2. present
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Chemistry: Spot Tests   back to top

927. Upper surface K reactivity     1. – (negative)     2. + deep yellow
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928. Upper surface C reactivity    ()  1. – ()
930. Upper surface KC reactivity    ()  1. – ()
931. Upper surface PD reactivity     1. –     8. + orange
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Index of all items of the project

In case that additional characters and states are required to be included in this data set, consult the LIAS Instructions to Participants and follow the procedures described there.

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