Botanische Staatssammlung München © 1995–2005
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Go to: Data maintenance � Nomenclature: Taxonomic Hierarchy � Nomenclature: Type Information � Nomenclature: Taxonomic Literature � Biogeography � Ecology � Lichen Photobionts � Thallus Morphology: General Morphology � Thallus Morphology: Thallus Size and Differentiation � Thallus Morphology: Outline � Thallus Morphology: Upper Thallus Surface � Thallus Morphology: Lower Thallus Surface � Thallus Anatomy: Upper Cortex � Reproduction � Ascocarp Morphology: General Ascocarp Morphology � Ascocarp Morphology: Margin � Ascocarp Morphology: Disk � Ascocarp Anatomy: Exciple � Ascocarp Anatomy: Epithecium � Ascocarp Anatomy: Hymenium � Ascocarp Anatomy: Subhymenium � Ascocarp Anatomy: Hypothecium � Ascocarp Anatomy: Asci � Ascocarp Anatomy: Ascospores � Conidioma Morphology � Conidioma Anatomy: Conidiogeneous Cells � Conidioma Anatomy: Conidia � Chemistry: Secondary Metabolites � Chemistry: Spot Tests � Index of all items of the project

Taxon name:

Data maintenance

1. Data set source [mandatory] [mandatory character] 2. compiled 3. template 5. standard item
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2. Data set author(s) [mandatory] [mandatory character] 42. Nash T.H. 52. Schramm R. & Nash T. H. None selected () ()

3. Data set reviewer(s) [mandatory character] 45. Nash T.H. 59. Schramm R. & Nash T.H.
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4. Data set revision status [mandatory character] 1. not revised 2. revised None selected () ()

5. Data set publication permission [mandatory character] 1. no 2. yes None selected () ()

6. Data set publication date

Nomenclature: Taxonomic Hierarchy

7. Current taxonomic status 1. accepted 2. basionymous
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8. Taxonomic rank [mandatory] [mandatory character] 2. species None selected () ()

11. Genus name [mandatory] [mandatory character] 343. Koerberia 494. Pannaria None selected () ()

12. Synonyms

13. Family name [mandatory] [mandatory character] 90. Pannariaceae Tuck. (1872) None selected () ()

14. Suborder name 6. Peltigerineae None selected () ()

15. Order name 9. Lecanorales None selected () ()

Nomenclature: Type Information

17. Type [mandatory character]

19. Protologue

20. Type specimen label data

Nomenclature: Taxonomic Literature

21. Taxonomic notes

22. Taxonomic literature

Biogeography

23. Hemisphere distribution () 1. Northern hemispheric ()

24. Floristic province {= zonal distribution} [mandatory] [mandatory character] 1. holarctic 2. arctic 3. subarctic 4. boreal 5. Mediterranean 6. subtropical 7. pantropical 8. neotropical 9. palaeotropic 10. subantarctic 11. antarctic 14. cosmopolitan
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25. Altitude [mandatory] [mandatory character] 1. coastal 2. colline 3. submontane 4. montane 5. upper montane 6. subalpine 7. alpine
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27. Continent distribution [mandatory] [mandatory character] 1. Africa 2. Antarctica 3. Asia-Temperate 4. Australasia 5. Europe 6. Northern America 7. Southern America 8. Pacific 9. Asia-Tropical
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28. Major geographic region 3. South-western Europe 6. Northern Africa 41. South-western U.S.A.
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29. European distribution records 19. France (excl. Corsica) 20. Portugal 26. Italy (incl. San Marino & The Vatican City, excl. Sicily, Sardinia)
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30. African distribution records () 4. Morocco ()

35. North American distribution records 46. Arizona (USA) 47. California (USA)
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38. Checklist records [mandatory] 13. Italy 15. Mexico 16. Morocco 20. Sonoran Desert 26. United States and Canada (continental)
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Ecology

43. Life habit () 2. biotroph (inhabiting living organisms) ()

44. Lifestyle [mandatory] [mandatory character] () 4. lichenized (= parasitic and / or mutual symbiotic with photobionts) ()

45. Substrate preference [mandatory character] 1. terricolous (on soil) 2. bryophytic (on bryophytes) 3. lignicolous (on wood) 4. corticolous (on bark) 6. saxicolous (on rock)
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47. Habitat insolation 1. sun-exposed 2. partially shaded 3. fully shaded
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48. Habitat moisture 1. dry 2. moist 3. wet
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53. Rock substrate mineral content 1. non-calciferous (lacking calcites) 2. calciferous (containing calcites)
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Lichen Photobionts

86. Primary photobiont genus 6. Calothrix 32. Myrmecia 35. Nostoc 42. Scytonema 43. Stichococcus 44. Stigonema
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91. Secondary photobiont presence 1. absent 2. present
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98. Secondary photobiont genus () 35. Nostoc ()

Thallus Morphology: General Morphology

104. Thallus growth form [mandatory character] 1. indistinct 2. crustose 3. foliose
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105. Thallus primary subdivision 1. continuous 2. divided
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107. Thallus ramification pattern 4. subdichotomous 5. irregular
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108. Crustose growth organization 1. leprose 2. granular 3. rimose 5. areolate (primarily areolate) 6. placodioid 7. squamulose
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109. Crustose growth structure 1. bullate 2. subsquamulose
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110. Crustose growth aggregation () 2. scattered (dispersed) ()

111. Crustose growth continuity 1. continuous throughout 2. dispersed along the periphery, continuous in the centre 3. dispersed throughout and distinctly seperate
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112. Foliose growth organization () 1. lobed ()

114. Foliose lobe outline 1. elongate 3. linear
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115. Foliose lobe separation 1. not laciniate 2. laciniate
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116. Foliose lobe orientation 1. horizontal 2. ascending 3. imbricate
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117. Foliose lobe front 1. angular 2. rounded
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121. Fruticose branch structure 1. terete 2. subterete 3. applanate (moderately flattened) 4. compressed (strongly flattened) 6. striate 7. ridged
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122. Fruticose branch flattening 1. bifacial 2. unifacial
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126. Thallus parts imbrication 1. not imbricate 2. imbricate
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128. Thallus parts distance [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					mm

130. Thallus parts length [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					mm long

132. Thallus parts width [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					mm wide

133. Thallus parts thickness [categorical] 1. thin 2. thick
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134. Thallus parts thickness [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					mm thick

Thallus Morphology: Thallus Size and Differentiation

151. Overall thallus diameter [categorical] 1. very small 2. small 3. medium-sized 4. large
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152. Overall thallus diameter [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					cm in diameter

157. Thallus apex width [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					mm wide

159. Thallus apex outline 1. rounded 2. truncate
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Thallus Morphology: Outline

160. Thallus perimeter outline 1. irregular 2. orbicular
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161. Thallus perimeter zonation 1. simple 2. zonate 3. lobed 4. arachnoid 5. fimbriate
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162. Thallus perimeter delimitation 1. indistinct 2. distinct
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164. Thallus margin structure 1. entire 3. dentate 4. laciniate 7. isidiate 8. phyllidiate 9. crenulate 10. dissected 11. lobulate 13. crenate
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167. Primal thallus differentiation 1. peripheral (forming a prothallus) 2. extending (forming a hypothallus)
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168. Primal thallus persistence 1. disappearing 2. persistent
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169. Primal thallus development 1. vestigial 2. pronounced 3. cushion forming
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170. Primal thallus colour 1. black 2. grey 4. green 6. brown 8. white
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Thallus Morphology: Upper Thallus Surface

171. Upper surface aspect 1. verrucose 2. verruculose
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172. Upper surface colour 4. grey 5. dark gray (= greyish black, blackish grey) 15. grey-green (= greyish green, greenish gray) 16. brownish green (= greenish brown, incl. olive, olivaceous) 17. bluish green (= greenish blue; incl. aeruginose, turquoise) 18. dark green (= blackish green, greenish black) 21. grey-blue (= bluish grey, greyish blue) 23. dark blue (= blackish blue, bluish black) 28. greenish yellow (= yellowish green, incl. lime green)
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174. Upper surface curvature 1. plane 2. subconcave 3. concave 4. subconvex 5. convex
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175. Upper surface glossiness 1. matt (opaque, dull) 2. glossy (shiny) 3. vitreous
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176. Upper surface roughness () 1. smooth ()

178. Upper surface special structures: presence 1. absent 2. present:
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196. Upper surface pruina: presence 1. epruinose 2. pruinose
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197. Upper surface pruina: abundance 1. scarce 2. abundant
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198. Upper surface pruina: colour 2. greyish 8. whitish
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230. Upper surface isidia: presence 1. absent 2. present
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231. Upper surface isidia: location () 2. laminal ()

232. Upper surface isidia: position {across the surface} 1. marginal 2. submarginal 3. apical
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235. Upper surface isidia: shape () 5. cylindrical ()

238. Upper surface isidia: ramification pattern 1. sparse 2. coralloid 3. moderate
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252. Upper surface soralia: presence 1. absent 2. present
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255. Upper surface soralia: colour 2. white 4. grey 30. brown
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256. Upper surface soralia: location 1. central 2. laminal 3. subperipheral 4. peripheral
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257. Upper surface soralia: position () 1. apical ()

259. Upper surface soralia: shape 1. semicapitate 2. capitate
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260. Upper surface soralia: outline () 1. erose (diffuse, unspecific) ()

262. Upper surface soralia: excavation () 2. fully excavated ()

264. Upper surface soralia: differentiation 1. not confluent 2. confluent (but not fusing)
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268. Upper surface soralia: development {from isidia} () 2. isidiate ()

269. Upper surface soredia: diameter [categorical] 1. grainy (coarse) 2. farinose (fine)
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275. Upper surface cephalodia: presence 1. absent 2. present
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278. Upper surface lobules: presence 1. absent 2. present
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Thallus Morphology: Lower Thallus Surface

282. Lower surface overall colour 2. white 4. grey 6. black 30. brown
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286. Lower surface attachment kind 1. by the whole lower surface 2. by holdfasts
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300. Lower surface rhizines: presence 1. absent 2. present (rhizinate)
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302. Lower surface rhizines: abundance 1. sparse 2. moderately abundant 3. abundant
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307. Lower surface rhizines: colour 4. grey 6. black 30. brown
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311. Lower surface rhizines: ramification pattern 1. irregular 2. dichotomous 3. squarrose 4. sparsely furcate
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326. Lower surface tomentum: presence 1. absent 2. present
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Thallus Anatomy: Upper Cortex

329. Upper cortex: presence 1. absent 2. present
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332. Upper cortex height [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm high

367. Upper cortex outer layer: structure 1. prosoplectenchymatous 2. paraplectenchymatous
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371. Upper cortex inner layer: structure 1. prosoplectenchymatous 2. paraplectenchymatous 3. scleroplectenchymatous
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Reproduction

414. Ascomata presence [mandatory] [mandatory character] 1. absent 2. present
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Ascocarp Morphology: General Ascocarp Morphology

421. Ascocarp location {on main thallus} 1. central 2. laminal 3. subperipheral 4. peripheral
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423. Ascocarp position {on primal thallus: pro- or hypothallus} 1. laminal 2. peripheral
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424. Ascocarp position {on primary thallus parts} 5. subterminal 6. terminal
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428. Ascocarp emergence () 1. not emerging (immersed, e.g., cryptolecanorine, cryptolecideine, aspicilioid) ()

429. Ascocarp attachment 1. adnate (semi-immersed) 2. sessile 3. substipitate
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431. Ascocarp abundance [categorical] 1. sparse 2. moderately abundant 3. abundant
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437. Ascocarp size [categorical] 1. small 2. large
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438. {orbicular} ascocarp diameter [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					mm in diam.

Ascocarp Morphology: Margin

447. Ascocarp margin: type 1. lecideine/biatorine 2. lecanorine (incl. zeorine)
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448. Ascocarp margin: structure 2. lobate 3. lobulate 4. crenulate
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449. Ascocarp margin: roughness () 1. smooth ()

451. Ascocarp margin: persistency 1. persistent 2. excluded
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452. Ascocarp margin: development 1. indistinct (thin) 2. distinct (moderately thickened) 3. prominent (thick)
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453. Ascocarp margin: colour 4. grey 6. black 13. green 30. brown
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463. Ascocarp margin: soredia 1. absent 2. present (sorediate)
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464. Ascocarp margin: isidia () 2. present (isidiate) ()

Ascocarp Morphology: Disk

488. Ascocarp disk: colour 6. black 7. red 13. green 24. yellow 30. brown
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491. Ascocarp disk: pruina presence 1. absent 2. present
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492. Ascocarp disk: pruina abundance 1. scarce 2. abundant
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493. Ascocarp disk: pruina colour 1. white 4. yellow
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Ascocarp Anatomy: Exciple

504. Ascocarp excipulum: pigment carbonization 1. biatorine (not carbonized) 2. lecideine (carbonized)
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Ascocarp Anatomy: Epithecium

627. Epithecium presence 1. absent 2. present
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629. Epithecium height [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm high

639. Epithecial hyphae apical cell pigmentation {in H2O} 10. brownish red (= reddish brown) 16. brownish green (= greenish brown, incl. olive, olivaceous) 17. bluish green (= greenish blue; incl. aeruginose, turquoise) 21. grey-blue (= bluish grey, greyish blue) 23. dark blue (= blackish blue, bluish black) 27. brownish yellow (= yellowish brown) 31. pale brown (= brownish white, whitish brown)
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641. Epithecial hyphae apical cell: pigment solubility {in 10% KOH} 1. insoluble in 10% KOH 2. soluble in 10% KOH
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642. Epithecial hyphae apical cell: pigment reactivity () 2. 10% KOH + ()

Ascocarp Anatomy: Hymenium

653. Hymenium height [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm high

654. Hymenium pigmentation {in H2O} 10. brownish red (= reddish brown) 27. brownish yellow (= yellowish brown) 31. pale brown (= brownish white, whitish brown)
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Ascocarp Anatomy: Subhymenium

684. Subhymenium height [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm high

699. Subhymenium pigmentation {in H2O} 2. white 4. grey 24. yellow 27. brownish yellow (= yellowish brown) 30. brown
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Ascocarp Anatomy: Hypothecium

714. Hypothecium plectenchyma structure 1. prosoplectenchymatous 2. paraplectenchymatous
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Ascocarp Anatomy: Asci

751. Ascus shape 1. globose 4. broadly clavate 5. clavate 6. narrowly clavate 7. obclavate 8. cylindrical
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753. Ascus length [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm long

754. Ascus width [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm wide

758. Ascus tholus iodine reactivity () 1. Lugol’s negative ()

760. Ascus tholus iodine reaction colour 1. + red 2. + light blue 3. + dark blue
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761. Ascus tholus amyloidy pattern 1. with amyloid tube 2. with amyloid cap 3. entirely amyloid throughout 5. with non-amyloid narrow axial body and amyloid convergent flanks towards apex
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Ascocarp Anatomy: Ascospores

784. Ascospore number [categorical] 2. c. 4 per ascus 3. c. 8 per ascus 4. 12-16 per ascus
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786. Ascospore shape 2. subglobose 3. oval 4. ellipsoid 5. broadly ellipsoid 16. fusiform
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787. Ascospore length [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm long

788. Ascospore width [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm wide

792. Ascospore septum presence 1. absent 2. present ()
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793. Ascospore septum orientation () 1. transversely septate ()

795. Ascospore {median} septum number [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					-transversally septate

Conidioma Morphology

816. Conidiomata presence [mandatory character] 1. absent 2. present
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817. Conidiomata kind () 1. pycnidial ()

818. Conidiomata abundance 1. sparse 2. moderately abundant 3. strongly abundant
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Conidioma Anatomy: Conidiogeneous Cells

865. Conidiogenous cells: position () 1. apical 2. intercalar
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Conidioma Anatomy: Conidia

873. Conidium shape 11. bacilliform 13. cylindrical
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879. Conidium length [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm long

880. Conidium width [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm wide

Chemistry: Secondary Metabolites

913. Known secondary metabolites: presence 1. absent 2. present
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915. Known secondary metabolites: name 231. atranorin 489. pannaric acid 491. pannarin
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916. Known secondary metabolites: substance class 2. (higher) aliphatic acids 14. (tri-)terpenoids
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917. Unknown secondary metabolites: presence 1. absent 2. present
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Chemistry: Spot Tests

927. Upper surface K reactivity 1. – (negative) 2. + deep yellow
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928. Upper surface C reactivity () 1. – ()

930. Upper surface KC reactivity () 1. – ()

931. Upper surface PD reactivity 1. – 8. + orange
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Index of all items of the project

In case that additional characters and states are required to be included in this data set, consult the LIAS Instructions to Participants and follow the procedures described there.

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