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Taxon name:

Data maintenance

1. Data set source [mandatory] [mandatory character] 3. template 5. standard item
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2. Data set author(s) [mandatory] [mandatory character] () ()

3. Data set reviewer(s) 1. N. N. 2. Aptroot A. 3. Arup U. 5. Beck A. 7. Calatayud V. 8. Carlin G. 9. Cáceres M. 11. Döbbeler P. 12. Döring H. 13. Egea J.M. & Torrente P. 14. Ekman S. 15. Elix J. A. 16. Esslinger T. 17. Feige G. B. 18. Feuerer T. 19. Gaya E. 20. Grube M. 21. Guderley R. 23. Helms G. 24. Hertel H. 25. Jørgensen P. M. 26. John V. 28. Kainz C. 29. Kärnefelt I. 30. Kasalicky T. 32. Keller C. 33. Kirschbaum U. 35. Koch A. 36. Kondratyuk S. 37. Lücking R. 38. Lumbsch H. T. 39. Lutzoni F. 40. Marbach B. 41. Mattsson J.-E. 42. Matzer M. & Mayrhofer H. 43. May P. 44. Moberg R. 45. Nash T.H. 46. Navarro-Rosinés P. 47. Nimis P. L. 48. Nordin A. 49. Owe-Larsson B. 50. Poelt J. 51. Printzen C. 52. Rambold G. 53. Randlane T. 54. Rico V. 55. Roux C. 56. Ryan B. 57. Scheidegger C. 58. Scholz P. () 59. Schramm R. & Nash T.H. 60. Schultz M. 61. Sheard J. 62. Sipman H. 63. Søchting U. 64. Staiger B. 65. Tehler A. 66. Thell A. 67. Tibell L. 68. Timdal E. 69. Triebel D. () 70. Verdon D. 71. Wedin M. 72. Zedda L.
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4. Data set revision status [mandatory character] 1. not revised 2. revised None selected () ()

5. Data set publication permission [mandatory character] 1. no 2. yes None selected () ()

6. Data set publication date

Nomenclature: Taxonomic Hierarchy

7. Current taxonomic status 1. accepted 2. basionymous
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8. Taxonomic rank [mandatory] [mandatory character] 1. genus None selected () ()

10. Taxon number

Lower range:	Mean:	Upper range:

11. Genus name [mandatory] [mandatory character] () ()

12. Synonyms

13. Family name [mandatory] [mandatory character] () ()

14. Suborder name () ()

15. Order name () ()

Nomenclature: Type Information

17. Type

Nomenclature: Taxonomic Literature

21. Taxonomic notes

22. Taxonomic literature
Alstrup V., Graphis Scripta 5: 96-104 (1993); Alstrup V. & Olech M., Fragm. Flor. Geobot. 41: 747-752 (1996); Calatayud V. & Triebel D., Nova Hedwigia 69(3-4): 439-448 (1999); Calatayud V. & Triebel D., Biblioth. Lichenol. 78: 27-33 (2001); Calatayud V. & Triebel D., Lichenologist 35(2): 103-116 (2003); Cole M.S. & Hawksworth D. L., Mycotaxon 77: 305-338 [330-332] (2001); Etayo J., Candollea 49: 245-249 (1994); Etayo J., Bull. Soc. Linn. Provençe 51: 153-162 [159-160] (2000); Etayo J. & Osorio H. S., Com. Bot. Mus. Nac. Hist. Nat. Antropol. 6(129): 1-19 [12-17] (2004); Hafellner J., Herzogia 10: 1-28 [23-24] (1994); Hafellner J., Bull. Soc. Linn. Provençe 45: 219-234 [230-232] (1994); Hafellner J. & Obermayer W., Cryptog. Bryol. Lichénol. 16: 177-190 (1995); Hafellner J., Triebel D., Ryan B.D. & Nash T.H., Mycotaxon 84: 293-329 (2002); Ihlen P.G., Lichenologist 30: 27 57 [53-54] (1998); Keissler K. von, Rabenh. Kryptog.-Fl. 8: 1-712 [365-366] (1930) - sub Pharcidia arthoniae; Kocourková J., Acta Mus. Nat. Pragae, Ser. B., Hist. Nat. 55(3-4): 59-169 [124-127] (2000); Lücking R., Cryptog. Mycol. 27: 121-147 [131-132] (2006); Martínez Moreno M.I., Ruzia 15: 1-200 [176-177] (1999); Matzer M., Mycol. Pap. 171: 1- 202 [155-169] (1996); Matzer M. & Hafellner J., Biblioth. Lichenol. 37: 1-138 [115-120](1990); Molitor F. & Diederich P., Bull. Soc. Naturalistes Luxemb. 98: 69- 92 [78-80] (1997); Roux C., Navarro Rosinés P. & Tranchida F., Cryptogamie, Bryol. Lichénol. 19(2-3): 221-228 (1998); Roux C. & Navarro Rosinés P., Bull. Soc. Linn. Provençe 45: 443-450 (1994); Renobales G., Guineana 2: 1-310 [172, 263] (1996); Roux C. & Triebel D., Bull. Soc. Linn. Provençe 45: 451-542 (1994); Roux C., Triebel D., Bricaud O. & Le Coeur D., Canad. J. Bot. 73: 622-672 (1995); Triebel D., Biblioth. Lichenol. 35: 1-278 [79-86] (1989)

Biogeography

38. Checklist records [mandatory] [mandatory character] 1. Australia 2. Austria 3. Bolivia 7. Germany 10. Great Britain 11. Guianas 13. Italy 17. New Guinea 18. New Zealand 20. Sonoran Desert 21. Sweden and Norway 22. Thailand 26. United States and Canada (continental) 28. Namibia 29. Republic of South Africa
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Ecology

43. Life habit 1. saprobic (= destruents) 2. biotroph (inhabiting living organisms)
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44. Lifestyle [mandatory] [mandatory character] 2. algicolous (= parasitic on eukaryotic algae) 3. phytopathogenic (= pathogens and/or parasites on plants) 4. lichenized (= parasitic and / or mutual symbiotic with photobionts) () 5. lichenicolous (= parasitic on lichens) 6. fungicolous (= parasitic on fungi)
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45. Substrate preference [mandatory character] 1. terricolous (on soil) 2. bryophytic (on bryophytes) 3. lignicolous (on wood) 4. corticolous (on bark) 5. epiphyllous (on living leaves)
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53. Rock substrate mineral content 1. non-calciferous (lacking calcites) 2. calciferous (containing calcites)
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Lichen Photobionts

79. Primary photobiont presence 1. absent 2. present
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80. Primary photobiont kind 1. cyanobacterial 2. chlorococcal 3. trentepohlioid
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86. Primary photobiont genus 1. Anabaena 2. Anacystis 4. Asterochloris 6. Calothrix 7. Cephaleuros 9. Chlorella 10. Chlorella saccharophila-group 11. Chlorosarcinopsis 12. Chroococcidiopsis 13. Chroococcus 15. Coccobotrys 16. Coccomyxa 18. Dichothrix 19. Dictyochloropsis 20. Dilabifilum 21. Elliptochloris 23. Gloeocapsa 24. Gloeocystis 26. Heterococcus 27. Hormathonema 28. Hyella 29. Hyphomorpha 30. Leptosira 31. Microcystis 32. Myrmecia 34. Nanochloris 35. Nostoc 36. Petroderma 37. Phycopeltis 39. Protococcus 40. Pseudochlorella 42. Scytonema 43. Stichococcus 44. Stigonema 45. Tolypothrix 46. Trebouxia 47. Trentepohlia 48. Trochiscia
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87. Primary photobiont family 1. Capsosiraceae 6. Chroococcaceae 9. Coccomyxaceae 11. Heterococcaceae 13. Hydrococcaceae 15. Lithodermataceae 16. Microchaetaceae 17. Microcystaceae 19. Nostocaceae 21. Pleurocapsaceae 23. Rivulariaceae 25. Scytonemataceae 26. Stigonemataceae 27. Trebouxiaceae 28. Trentepohliaceae 29. Xenococcaceae 30. genus incertae sedis
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88. Primary photobiont order 3. Chlorococcales 5. Chroococcales 7. Dictyochloropsis-group 8. Ectocarpales 10. Leptosira-group 11. Nostocales 14. Stigonematales 15. Trebouxiales 16. Trentepohliales 17. Tribonematales 20. family of unknown placement (incertae sedis)
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89. Primary photobiont class 1. Cyanobacteria 2. Xanthophyceae 3. Phaeophyceae 4. Chlorophyceae 5. Trebouxiophyceae 7. Ulvophyceae H. order of unknown placement (incertae sedis)
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90. Primary photobiont kingdom 1. Prokaryota 2. Eukariota
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91. Secondary photobiont presence 1. absent 2. present
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98. Secondary photobiont genus 1. Anabaena 2. Anacystis 3. Aphanocapsa 23. Gloeocapsa 35. Nostoc 42. Scytonema 44. Stigonema
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99. Secondary photobiont family 7. Chroococcaceae 18. Microcystaceae 20. Nostocaceae 26. Scytonemataceae 27. Stigonemataceae
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100. Secondary photobiont order 5. Chroococcales 11. Nostocales 14. Stigonematales
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101. Secondary photobiont class () 1. Cyanobacteria ()

102. Secondary photobiont kingdom () 1. Prokaryota ()

Thallus Morphology: General Morphology

104. Thallus growth form [mandatory character] 1. indistinct 2. crustose 3. foliose 4. fruticose
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105. Thallus primary subdivision () 1. continuous ()

108. Crustose growth organization 1. leprose 2. granular 3. rimose 5. areolate (primarily areolate) 6. placodioid 7. squamulose 8. subfruticose (minutely fruticose, dactyliform) 9. filamentose
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109. Crustose growth structure () 4. peltate ()

112. Foliose growth organization 1. lobed 2. umbilicate
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Thallus Morphology: Outline

168. Primal thallus persistence 1. disappearing 2. persistent
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Thallus Morphology: Upper Thallus Surface

172. Upper surface colour 2. white 4. grey 6. black 7. red 8. pale red (= reddish white, whitish red, incl. pink, rosé) 13. green 16. brownish green (= greenish brown, incl. olive, olivaceous) 24. yellow 26. grey-yellow (= greyish yellow, yellowish grey) 27. brownish yellow (= yellowish brown) 30. brown 34. orange (= reddish yellow, yellowish red)
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178. Upper surface special structures: presence 1. absent 2. present:
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201. Upper surface pseudocyphellae: presence 1. absent 2. present
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218. Upper surface cilia: presence 1. eciliate 2. ciliate
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225. Upper surface hairs: presence 1. absent 2. present
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230. Upper surface isidia: presence 1. absent 2. present
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252. Upper surface soralia: presence 1. absent 2. present
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275. Upper surface cephalodia: presence 1. absent 2. present
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278. Upper surface lobules: presence 1. absent 2. present
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279. Upper surface granules: presence 1. absent 2. present
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280. Upper surface thalloconidia: presence 1. absent 2. present
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Thallus Morphology: Lower Thallus Surface

286. Lower surface attachment kind 1. by the whole lower surface 2. by holdfasts 3. lacking
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287. Lower surface special structures: presence 1. absent 2. present:
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288. Lower surface pseudocyphellae: presence 1. absent 2. present
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295. Lower surface cyphellae: presence 1. absent 2. present
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300. Lower surface rhizines: presence 1. absent 2. present (rhizinate)
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319. Lower surface thalloconidia: presence 1. absent 2. present
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325. Lower surface cavernae: presence 1. absent 2. present
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326. Lower surface tomentum: presence 1. absent 2. present
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Thallus Anatomy: Medulla

393. Medulla euamyloid reactivity 1. in Lugol's solution negative 2. in Lugol's solution positive
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394. Medulla hemiamyloid reactivity 1. not hemiamyloid 2. hemiamyloid
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Reproduction

414. Ascomata presence [mandatory] [mandatory character] 1. absent 2. present
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Ascocarp Morphology: General Ascocarp Morphology

418. Ascocarp structure [mandatory character] 1. apothecioid 2. perithecioid 3. cleistothecioid
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420. Ascocarp outline 1. orbicular 2. lingulate 3. irregular 4. linear 5. stellate
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421. Ascocarp location {on main thallus} 2. laminal 4. peripheral
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424. Ascocarp position {on primary thallus parts} () 6. terminal ()

428. Ascocarp emergence () 1. not emerging (immersed, e.g., cryptolecanorine, cryptolecideine, aspicilioid) ()

429. Ascocarp attachment 1. adnate (semi-immersed) 2. sessile 3. substipitate 4. stipitate
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436. Ascocarp ascostroma formation () 3. stromatic ()

Ascocarp Morphology: Wall

443. Ascocarp wall: carbonization 1. not carbonized 2. carbonized only in parts 3. carbonized around the ostiole, lacking a clypeus 4. with a carbonized black clypeus around the ostiole
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444. Ascocarp wall: fusion {of neighboured perithecia} 1. absent 2. present
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Ascocarp Morphology: Margin

452. Ascocarp margin: development 1. indistinct (thin) 2. distinct (moderately thickened) 3. prominent (thick)
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468. Ascocarp margin: external filaments presence 1. absent 2. present
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Ascocarp Anatomy: Exciple

505. Ascocarp excipulum: layer pigmentation 2. white 4. grey 6. black 7. red 8. pale red (= reddish white, whitish red, incl. pink, rosé) 11. bluish red (= reddish blue, incl. violet, purple) 13. green 16. brownish green (= greenish brown, incl. olive, olivaceous) 24. yellow 27. brownish yellow (= yellowish brown) 30. brown 34. orange (= reddish yellow, yellowish red)
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Ascocarp Anatomy: Periphyses

613. Periphyses presence 1. absent 2. present
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Ascocarp Anatomy: Epithecium

639. Epithecial hyphae apical cell pigmentation {in H2O} 1. hyaline (translucent) 4. grey 6. black 7. red 8. pale red (= reddish white, whitish red, incl. pink, rosé) 11. bluish red (= reddish blue, incl. violet, purple) 13. green 16. brownish green (= greenish brown, incl. olive, olivaceous) 24. yellow 27. brownish yellow (= yellowish brown) 30. brown 34. orange (= reddish yellow, yellowish red)
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Ascocarp Anatomy: Hymenium

650. Hymenium persistence () 2. mazaedioid (asci soon disintregating) ()

661. Hymenium inspersion: type () 2. inspersed with oil droplets ()

667. Hymenium euamyloid reaction 1. Lugol’s negative 2. Lugol’s positive
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668. Hymenium hemiamyloid reaction 1. not hemiamyloid 2. hemiamyloid
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Ascocarp Anatomy: Interascal Hyphae

673. Interascal hyphae presence 1. absent 2. present
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676. Interascal hyphae ramification 1. scarcely branched 2. distinctly branched
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677. Interascal hyphae anastomoses 1. not or scarcely anastomosed 2. distinctly anastomosed
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Ascocarp Anatomy: Hypothecium

708. Hypothecium pigmentation {in H2O} 2. white 4. grey 6. black 7. red 8. pale red (= reddish white, whitish red, incl. pink, rosé) 11. bluish red (= reddish blue, incl. violet, purple) 13. green 16. brownish green (= greenish brown, incl. olive, olivaceous) 24. yellow 27. brownish yellow (= yellowish brown) 30. brown 34. orange (= reddish yellow, yellowish red)
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Ascocarp Anatomy: Asci

757. Ascus tholus thickness 1. not thickened 2. thickened
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758. Ascus tholus iodine reactivity 1. Lugol’s negative 2. Lugol’s positive
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761. Ascus tholus amyloidy pattern 1. with amyloid tube 2. with amyloid cap 3. entirely amyloid throughout 4. with amyloid cap or tube and flanks 5. with non-amyloid narrow axial body and amyloid convergent flanks towards apex 6. with non-amyloid wide axial body and divergent amyloid flanks towards apex
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768. Ascus ocular chamber development 1. indistinct 3. broad
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769. Ascus dehiscence type 1. prototunicate 2. lecanoralean 3. unitunicate 4. verrucarial 5. bitunicate
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778. Exoascus {euamyloid} reactivity 1. in in Lugol's Solution Solution negative 2. in in Lugol's Solution Solution positive
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779. Exoascus {hemiamyloid} reactivity 1. not hemiamyloid 2. hemiamyloid
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Ascocarp Anatomy: Ascospores

784. Ascospore number [categorical] 1. 1–2 per ascus 2. c. 4 per ascus 3. c. 8 per ascus 4. 12-16 per ascus 5. 16-32 per ascus 6. more than 32 per ascus
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786. Ascospore shape 1. globose 4. ellipsoid 5. broadly ellipsoid 6. ovoid 10. oblong-obtuse 11. oblong 13. cylindrical 16. fusiform 17. filiform 23. clavate 28. semilunate 31. reniform (incl. fabiform) 32. curved 35. allantoid 39. irregular
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787. Ascospore length [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm long

788. Ascospore width [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm wide

790. Ascospore attenuation 1. obtuse 2. aciculate
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792. Ascospore septum presence 1. absent 2. present
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793. Ascospore septum orientation 1. transversely septate 2. muriform
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795. Ascospore {median} septum number [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					-transversally septate

797. Ascospore septum ultrastructure 1. proper wall septum (omniseptate) 2. endospore septum (endoseptate)
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802. Ascospore wall width [categorical] 1. thin 2. thick
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804. Ascospore wall differentiation {visible in the light microscope} () 2. distinctly differentiated (into primary and secondary wall) ()

807. {septal} ascospore wall thickening 1. absent 2. present
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808. Ascospore wall lateral septal constrictions 1. absent 2. present
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810. Ascospore wall pigmentation 1. hyaline (translucent) 2. grey 4. pale brown (= brownish white, whitish brown) 5. middle brown 7. dark brown (= brownish black, blackish brown)
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811. Ascospore wall iodine reactivity {euamyloid} 1. in Lugol's Solution negative 2. in Lugol's Solution positive
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812. Ascospore perispore ornamentation 1. not ornamented (psilate, smooth) 2. ornamented
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Conidioma Morphology

816. Conidiomata presence [mandatory character] 1. absent 2. present
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817. Conidiomata kind 1. pycnidial 2. hyphomycetous 3. hyphophorous 4. campylidial
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819. Conidiomata exposure 1. immersed 2. adnate (semi-immersed) 3. sessile 4. stipitate
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826. Conidiomata location 2. laminal (on thallus) 4. peripheral (on thallus)
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Conidioma Anatomy: Conidiogeneous Cells

865. Conidiogenous cells: position 1. apical 2. intercalar
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Conidioma Anatomy: Conidia

873. Conidium shape 1. globose 4. ellipsoid 11. bacilliform 16. fusiform 17. filiform 22. lageniform 32. reniform 33. curved 37. bifusiform
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874. Conidium size [categorical] 1. microconidial 3. macroconidial
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875. Conidium ramification: presence 1. absent 2. present
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882. Conidium septation: presence 1. absent 2. present
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884. {transversal} conidium septation [numerical]

Lower range:	Upper range:	Unit:
		-septate

886. Conidium cell wall pigmentation () 1. hyaline (translucent) ()

Chemistry: Secondary Metabolites

913. Known secondary metabolites: presence 1. absent 2. present
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916. Known secondary metabolites: substance class 1. amino acid derivatives 2. (higher) aliphatic acids 3. (anthra-)quinones 4. benzyl esters 6. dibenzofurans [and usnic acids] 7. diphenyl ethers 8. orcinol depsides 9. orcinol depsidones 10. ß-orcinol depsides [incl. atranorin] 11. ß-orcinol depsidones 12. pulvinic acid derivatives 13. secalonic acids 14. (tri-)terpenoids 15. xanthones
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