Botanische Staatssammlung München © 1995–2005
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Taxon name:

Data maintenance

1. Data set source [mandatory] [mandatory character] 1. authentic 2. compiled 3. template 4. checklist item 5. standard item
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2. Data set author(s) [mandatory] [mandatory character] 42. Nash T.H. 52. Schramm R. & Nash T. H. None selected () ()

3. Data set reviewer(s) [mandatory character] () 59. Schramm R. & Nash T.H. ()

4. Data set revision status [mandatory character] 1. not revised 2. revised None selected () ()

5. Data set publication permission [mandatory character] 1. no 2. yes None selected () ()

6. Data set publication date

Nomenclature: Taxonomic Hierarchy

7. Current taxonomic status () 1. accepted ()

8. Taxonomic rank [mandatory] [mandatory character] 2. species 3. subspecies 4. variety 5. form None selected () ()

10. Taxon number

Mean:

11. Genus name [mandatory] [mandatory character] () ()

12. Synonyms

13. Family name [mandatory] [mandatory character] 3. Acarosporaceae Zahlbr. (1906) None selected () ()

14. Suborder name 2. Acarosporineae None selected () ()

15. Order name 9. Lecanorales None selected () ()

Nomenclature: Type Information

17. Type [mandatory character]

19. Protologue

20. Type specimen label data

Nomenclature: Taxonomic Literature

21. Taxonomic notes

22. Taxonomic literature

Biogeography

23. Hemisphere distribution () 1. Northern hemispheric ()

24. Floristic province {= zonal distribution} [mandatory] [mandatory character] 1. holarctic 2. arctic 3. subarctic 4. boreal 5. Mediterranean 6. subtropical 7. pantropical 8. neotropical 9. palaeotropic 10. subantarctic 11. antarctic 14. cosmopolitan
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25. Altitude [mandatory] [mandatory character] 1. coastal 2. colline 3. submontane 4. montane 5. upper montane 6. subalpine 7. alpine
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27. Continent distribution [mandatory] [mandatory character] 1. Africa 2. Antarctica 3. Asia-Temperate 4. Australasia 5. Europe 6. Northern America 7. Southern America 8. Pacific 9. Asia-Tropical
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38. Checklist records [mandatory] 15. Mexico 20. Sonoran Desert 26. United States and Canada (continental)
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Ecology

43. Life habit () 2. biotroph (inhabiting living organisms) ()

44. Lifestyle [mandatory] [mandatory character] 4. lichenized (= parasitic and / or mutual symbiotic with photobionts) 5. lichenicolous (= parasitic on lichens)
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45. Substrate preference [mandatory character] 1. terricolous (on soil) 2. bryophytic (on bryophytes) 3. lignicolous (on wood) 4. corticolous (on bark) 6. saxicolous (on rock)
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46. Thallus-substrate interaction 1. endosubstratic 3. episubstratic
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47. Habitat insolation 1. sun-exposed 2. partially shaded 3. fully shaded
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48. Habitat moisture 1. dry 2. moist 3. wet
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53. Rock substrate mineral content 1. non-calciferous (lacking calcites) 2. calciferous (containing calcites)
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Lichen Photobionts

79. Primary photobiont presence 1. absent 2. present
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80. Primary photobiont kind () 2. chlorococcal ()

87. Primary photobiont family () 27. Trebouxiaceae ()

91. Secondary photobiont presence () 1. absent ()

Thallus Morphology: General Morphology

104. Thallus growth form [mandatory character] 1. indistinct 2. crustose 3. foliose
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105. Thallus primary subdivision 1. continuous 2. divided
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108. Crustose growth organization 2. granular 3. rimose 5. areolate (primarily areolate) 6. placodioid 7. squamulose 8. subfruticose (minutely fruticose, dactyliform)
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109. Crustose growth structure 1. bullate 2. subsquamulose
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110. Crustose growth aggregation () 2. scattered (dispersed) ()

112. Foliose growth organization 1. lobed 2. umbilicate
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114. Foliose lobe outline () 1. elongate ()

117. Foliose lobe front 1. angular 2. rounded
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128. Thallus parts distance [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					mm

129. Thallus parts length [categorical] 1. short 2. long
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130. Thallus parts length [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					mm long

131. Thallus parts width [categorical] 1. narrow 2. wide
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132. Thallus parts width [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					mm wide

133. Thallus parts thickness [categorical] 1. thin 2. thick
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134. Thallus parts thickness [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					mm thick

Thallus Morphology: Thallus Size and Differentiation

151. Overall thallus diameter [categorical] 1. very small 2. small 3. medium-sized
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152. Overall thallus diameter [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					cm in diameter

154. Thallus joints main branch {basal} constriction 1. basally not constricted 2. basally constricted
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155. Thallus joints secondary branch {basal} constriction 1. basally not constricted 2. basally constricted
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157. Thallus apex width [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					mm wide

159. Thallus apex outline 1. rounded 2. truncate
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Thallus Morphology: Outline

160. Thallus perimeter outline 1. irregular 2. orbicular
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161. Thallus perimeter zonation 1. simple 2. zonate 3. lobed 4. arachnoid 5. fimbriate
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164. Thallus margin structure 1. entire 3. dentate 4. laciniate 6. undulate 9. crenulate 11. lobulate 13. crenate
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Thallus Morphology: Upper Thallus Surface

171. Upper surface aspect 1. verrucose 2. verruculose
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172. Upper surface colour 2. white 24. yellow 28. greenish yellow (= yellowish green, incl. lime green) 30. brown 31. pale brown (= brownish white, whitish brown)
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174. Upper surface curvature 1. plane 2. subconcave 3. concave 4. subconvex 5. convex
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175. Upper surface glossiness 1. matt (opaque, dull) 2. glossy (shiny) 3. vitreous
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176. Upper surface roughness 1. smooth 2. rough
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177. Upper surface texture 3. byssoid 4. erose
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196. Upper surface pruina: presence 1. epruinose 2. pruinose
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197. Upper surface pruina: abundance 1. scarce 2. abundant
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198. Upper surface pruina: colour 2. greyish 8. whitish
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199. Upper surface pruina: distribution 1. continuous 2. patchy
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200. Upper surface pruina: location 1. central 3. peripheral
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230. Upper surface isidia: presence 1. absent 2. present
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252. Upper surface soralia: presence 1. absent 2. present
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Thallus Morphology: Lower Thallus Surface

281. Lower surface presence 1. absent 2. present
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282. Lower surface overall colour 10. brownish red (= reddish brown) 16. brownish green (= greenish brown, incl. olive, olivaceous) 27. brownish yellow (= yellowish brown) 31. pale brown (= brownish white, whitish brown) 32. grey-brown (= greyish brown, brownish grey)
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Thallus Anatomy: Upper Cortex

332. Upper cortex height [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm high

334. Upper cortex plectenchyma structure 1. prosoplectenchymatous 2. paraplectenchymatous
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339. Upper cortex hyphal cell structure () 1. chondroid ()

Thallus Anatomy: Medulla

380. Medulla height [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm thick

384. Medulla density 1. compact 2. dense 3. loose 4. very loose, becoming hollow
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393. Medulla euamyloid reactivity 1. in Lugol's solution negative 2. in Lugol's solution positive
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Thallus Anatomy: Lower Cortex

397. Lower cortex presence 1. absent 2. present
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Reproduction

414. Ascomata presence [mandatory] [mandatory character] 1. absent 2. present
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Ascocarp Morphology: General Ascocarp Morphology

418. Ascocarp structure [mandatory character] 1. apothecioid 2. perithecioid
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428. Ascocarp emergence () 1. not emerging (immersed, e.g., cryptolecanorine, cryptolecideine, aspicilioid) ()

429. Ascocarp attachment 1. adnate (semi-immersed) 2. sessile 3. substipitate
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432. Ascocarp abundance [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					per thallus

437. Ascocarp size [categorical] 1. small 2. large
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438. {orbicular} ascocarp diameter [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					mm in diam.

Ascocarp Morphology: Margin

447. Ascocarp margin: type 1. lecideine/biatorine 2. lecanorine (incl. zeorine)
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449. Ascocarp margin: roughness 1. smooth 5. rough
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457. Ascocarp margin: width [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm wide

Ascocarp Morphology: Disk

487. Ascocarp disk: curvature 1. urceolate (strongly concave) 2. excavate (concave) 3. plane (expanded) 4. weakly convex
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488. Ascocarp disk: colour 24. yellow 27. brownish yellow (= yellowish brown) 30. brown 32. grey-brown (= greyish brown, brownish grey) 33. dark brown (= brownish black, blackish brown)
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489. Ascocarp disk: exposure 1. not exposed 2. partly exposed 3. fully exposed
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491. Ascocarp disk: pruina presence 1. absent 2. present
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Ascocarp Anatomy: Exciple

497. Ascocarp excipulum: plectenchyma structure () 2. paraplectenchymatous ()

498. Ascocarp excipulum: hyphal orientation () 3. not easily discerned (strongly gelatinized or carbonized) ()

504. Ascocarp excipulum: pigment carbonization 1. biatorine (not carbonized) 2. lecideine (carbonized)
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Ascocarp Anatomy: General Amphithecium Anatomy

559. Amphithecial photobiont abundance 1. absent 2. sparse (eulecanorine) 3. abundant
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Ascocarp Anatomy: Episamma

622. Episamma presence 1. absent 2. present
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Ascocarp Anatomy: Epithecium

627. Epithecium presence 1. absent 2. present
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629. Epithecium height [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm high

630. Epithecial hyphae apical cell: shape 1. simple 2. spathulate 3. subulate
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633. Epithecial hyphae apical cell: width [categorical] 1. not swollen 2. slightly swollen 3. distinctly swollen
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634. Epithecial hyphae apical cell: width [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm wide

639. Epithecial hyphae apical cell pigmentation {in H2O} 1. hyaline (translucent) 10. brownish red (= reddish brown) 16. brownish green (= greenish brown, incl. olive, olivaceous) 26. grey-yellow (= greyish yellow, yellowish grey) 27. brownish yellow (= yellowish brown) 28. greenish yellow (= yellowish green, incl. lime green) 31. pale brown (= brownish white, whitish brown) 33. dark brown (= brownish black, blackish brown)
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641. Epithecial hyphae apical cell: pigment solubility {in 10% KOH} 1. insoluble in 10% KOH 2. soluble in 10% KOH
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642. Epithecial hyphae apical cell: pigment reactivity () 2. 10% KOH + ()

645. Epithecial hyphae subapical cell width [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm wide

Ascocarp Anatomy: Hymenium

653. Hymenium height [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm high

654. Hymenium pigmentation {in H2O} 2. white 3. pale grey (= greyish white, whitish grey) 14. pale green (= greenish white, whitish green) 25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine) 31. pale brown (= brownish white, whitish brown)
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661. Hymenium inspersion: type 1. not inspers 2. inspersed with oil droplets 3. inspersed with crystals
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Ascocarp Anatomy: Interascal Hyphae

676. Interascal hyphae ramification 1. scarcely branched 2. distinctly branched
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679. Interascal hyphae cell length [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm long

Ascocarp Anatomy: Subhymenium

684. Subhymenium height [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm high

699. Subhymenium pigmentation {in H2O} 2. white 3. pale grey (= greyish white, whitish grey) 25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine) 31. pale brown (= brownish white, whitish brown)
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Ascocarp Anatomy: Hypothecium

707. Hypothecium height [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm high

708. Hypothecium pigmentation {in H2O} 2. white 3. pale grey (= greyish white, whitish grey) 25. pale yellow (= whitish yellow, yellowish white, incl. lemon, citrine) 31. pale brown (= brownish white, whitish brown)
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716. Hypothecium prosoplectenchyma type 2. serioplectenchymatous 9. dense textura intricata
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717. Hypothecium hyphal orientation 2. anticlinal 3. irregular
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Ascocarp Anatomy: Asci

751. Ascus shape 1. globose 4. broadly clavate 5. clavate 6. narrowly clavate 7. obclavate 8. cylindrical
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753. Ascus length [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm long

754. Ascus width [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm wide

758. Ascus tholus iodine reactivity 1. Lugol’s negative 2. Lugol’s positive
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761. Ascus tholus amyloidy pattern () 3. entirely amyloid throughout ()

769. Ascus dehiscence type () 2. lecanoralean ()

Ascocarp Anatomy: Ascospores

784. Ascospore number [categorical] 5. 16-32 per ascus 6. more than 32 per ascus
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786. Ascospore shape 1. globose 2. subglobose 3. oval 4. ellipsoid 5. broadly ellipsoid
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787. Ascospore length [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm long

788. Ascospore width [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm wide

792. Ascospore septum presence 1. absent 2. present
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810. Ascospore wall pigmentation () 1. hyaline (translucent) ()

Conidioma Morphology

817. Conidiomata kind () 1. pycnidial ()

Conidioma Anatomy: Pycnidia

833. Pycnidium shape 1. globular 2. subglobular 4. ovoid 6. pyriform
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Conidioma Anatomy: Conidia

873. Conidium shape 1. globose 2. subglobose 4. ellipsoid 5. broadly ellipsoid
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879. Conidium length [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm long

880. Conidium width [numerical]

Min:	Lower range:	Mean:	Upper range:	Max:	Unit:
					µm wide

Chemistry: Secondary Metabolites

913. Known secondary metabolites: presence 1. absent 2. present
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915. Known secondary metabolites: name 336. gyrophoric acid 470. norstictic acid 530. rhizocarpic acid 596. vulpinic acid
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916. Known secondary metabolites: substance class 8. orcinol depsides 9. orcinol depsidones 12. pulvinic acid derivatives
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917. Unknown secondary metabolites: presence 1. absent 2. present
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Chemistry: Spot Tests

927. Upper surface K reactivity () 1. – (negative) ()

928. Upper surface C reactivity 1. – 5. + red
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930. Upper surface KC reactivity 1. – 5. + red
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931. Upper surface PD reactivity () 1. – ()

937. Medulla K reactivity 1. – 8. + orange
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938. Medulla C reactivity 1. – 5. + red
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940. Medulla KC reactivity () 5. + red ()

941. Medulla PD reactivity 1. – 5. + red
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